During the late Oligocene (23 mya) the New Zealand landmass was reduced to approximately 18% of its current area. It has been hypothesized that this event, known as the Oligocene Drowning, caused population bottlenecking and mass extinction. Using phylogenetic methods, we examine the effect of this and other environmental events on the hyper‐diverse Zopheridae beetles (162 morphospecies), which largely inhabit leaf litter and dead wood.

中文翻译：

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在新西兰渐新世溺水中幸存了多种多样的超大型Zopheridae甲虫

在渐新世晚期（23 mya），新西兰的陆地减少到其当前面积的约18％。据推测，这一事件被称为渐新世溺水，造成人口瓶颈和大规模灭绝。使用系统发育方法，我们研究了此事件和其他环境事件对主要栖息于枯枝落叶和枯木的超多样性Zopheridae甲虫（162个形态物种）的影响。