In light of the available discoveries in the field, this review manuscript discusses on plant reproduction mechanism and molecular players involved in the process. Sperm cells in angiosperms are immotile and are physically distant to the female gametophytes (FG). To secure the production of the next generation, plants have devised a clever approach by which the two sperm cells in each pollen are safely delivered to the female gametophyte where two fertilization events occur (by each sperm cell fertilizing an egg cell and central cell) to give rise to embryo and endosperm. Each of the successfully fertilized ovules later develops into a seed. Sets of macromolecules play roles in pollen tube (PT) guidance, from the stigma, through the transmitting tract and funiculus to the micropylar end of the ovule. Other sets of genetic players are involved in PT reception and in its rupture after it enters the ovule, and yet other sets of genes function in gametic fusion. Angiosperms have come long way from primitive reproductive structure development to today's sophisticated, diverse, and in most cases flamboyant organ. In this review, we will be discussing on the intricate yet complex molecular mechanism of double fertilization and how it might have been shaped by the evolutionary forces focusing particularly on the model plant Arabidopsis.

中文翻译：

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开花植物的受精：精子细胞传递的奥德赛。

鉴于该领域的现有发现，本综述手稿讨论了植物繁殖机制和参与该过程的分子参与者。被子植物中的精子细胞是不动的，并且与雌性配子体 (FG) 物理距离较远。为了确保下一代的生产，植物设计了一种聪明的方法，通过这种方法，每个花粉中的两个精子细胞被安全地输送到雌配子体，在那里发生两次受精事件（每个精子细胞与一个卵细胞和中央细胞受精）产生胚和胚乳。每个成功受精的胚珠后来都发育成种子。成组的大分子在花粉管 (PT) 引导中发挥作用，从柱头，通过传输道和绳索到胚珠的珠孔端。其他基因组参与了 PT 接收和它进入胚珠后的破裂，还有其他基因组在配子融合中起作用。被子植物从原始的生殖结构发展到今天的复杂、多样化，并且在大多数情况下是华丽的器官，已经走过了漫长的道路。在这篇综述中，我们将讨论复杂而复杂的双重受精分子机制，以及特别关注模式植物拟南芥的进化力量如何塑造它。