Freshwater gobies played an important role in the Miocene paleolakes of central and southeastern Europe. Much data have been gathered from isolated otoliths, but articulated skeletons are relatively rare. Here, we review a rich assemblage of articulated gobies with abundant otoliths in situ from the late early Miocene lake deposits of Klinci in the Valjevo freshwater lake of the Valjevo-Mionica Basin of Serbia. The fauna was originally described by Anđelković in 1978, who noted many different fishes, including one goby (Gobius multipinnatus H. v. Meyer 1848), and was subsequently revised by Gaudant (1998), who collapsed all previously recognized species into a single gobiid species that he described as new, namely Gobius serbiensis Gaudant 1998. Our review resulted in the recognition of three highly adaptive extinct freshwater gobiid genera with four species being divided among them: Klincigobius andjelkovicae n.gen. and n.sp., Klincigobius serbiensis (Gaudant 1998), Rhamphogobius varidens n.gen. and n.sp., and Toxopyge campylus n.gen. and n.sp. Otoliths were found in situ in all four species, which allowed for the allocation of multiple previously described otolith-based species to these extinct gobiid genera. With the help of correlations among isolated otoliths, it is now possible to outline putative paleobioprovinces of fishes in the early Miocene freshwater lakes of central and southeastern Europe. The gobiid freshwater fauna from the early to early middle Miocene of Europe was found to not be related to any of the extant European freshwater or brackish water gobies. Instead, they likely represented an earlier cycle of freshwater adaptation in a warm, subtropical environment that vanished after the Miocene Climate Optimum at c. 14 Ma, when it gave way to a new phase of freshwater goby adaptation that led to the present day Ponto-Caspian goby stock.

中文翻译：

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从克林奇（塞尔维亚）的中新世早期消失的淡水虾虎鱼鱼类动物群（Teleostei，Gobiidae）

淡水虾虎鱼在中欧和东南欧的中新世古湖中起着重要作用。已经从孤立的耳石中收集了许多数据，但是铰接的骨骼相对较少。在这里，我们从塞尔维亚Valjevo-Mionica盆地的Valjevo淡水湖中的Klinci早期中新世晚期晚期沉积岩中，总结了带有大量耳石的多臂虎虾的丰富组合。该动物群最初是由Anđelković于1978年描述的，他注意到许多不同的鱼类，其中包括一种虾虎鱼（Gobius multipinnatus H. v。Meyer 1848），随后由Gaudant（1998）进行了修订，后者将所有先前公认的物种折叠为一个戈壁他将其描述为新物种，即Gobius serbiensis Gaudant 1998。我们的审查导致对三个高度适应的灭绝淡水戈壁鱼属的认识，其中四个物种被划分为：Klincigobius和jelkovicae n.gen。和N.sp.，Klincigobius serbiensis（Gaudant 1998），Rhamphogobius varidens n.gen。n.sp.和Toxopyge campylus n.gen。和n.sp. 在所有四个物种的原位都发现了耳石，这允许将多个先前描述的基于耳石的物种分配给这些灭绝的戈壁动物属。借助于孤立的耳石之间的相关性，现在有可能勾勒出中欧和东南欧中新世早期淡水湖中鱼类的假定古生物区。发现欧洲中新世早期至中中期的戈壁水生动物区系与任何现存的欧洲淡水或淡水虾虎鱼都不相关。代替，它们可能代表了在温暖的亚热带环境中淡水适应的早期周期，该环境在中新世气候最优化于1989年后消失。14 Ma，当它进入淡水虾虎鱼适应的新阶段时，导致了今天的里海虾虎鱼种群。