A new fossil meso-flora consisting of dispersed cuticles has been discovered within the Upper Member of the Middle Triassic (late Anisian) Mukheiris Formation at the eastern rim of the Dead Sea in Jordan. The assemblage consists of a number of different morpho-types of cuticles which can all be assigned to gymnosperms. Cuticles which can be identified as the corystospermalean taxon Dicroidium are fairly abundant and clearly dominate the assemblage, whereas cuticles assignable to other gymnosperms are rather rare. As so far all Triassic records of Dicroidium come from localities at palaeolatitudes above 35°S, these cuticles represent the first record of this genus within the Triassic palaeotropics. Our analysis of this meso-flora suggests that during the late Anisian, as in other regions worldwide, a gymnosperm component of the terrestrial vegetation on the Arabian Peninsula was present. Compared to the palynoflora, which yielded various types of conifer pollen, and pollen of other gymnosperm groups, as well as abundant spores of ferns and lycophytes, the meso-flora is so far lacking a number of plant groups which were present in the source vegetation. It is likely that the differences can be explained by taphonomical biases. The data from cuticles and palynology together prove that either regional terrestrial ecosystems had already recovered, at least to some extent, from the devastating effects of the end-Permian ecological crisis or that these effects were less devastating, at least with regard to vegetation, than assumed by previous studies.

中文翻译：

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基于约旦Anisian Mukheiris地层的分散角质层，三叠纪古热带中的Dicroidium的第一个记录

在约旦死海东缘的中三叠世（晚安尼期）Mukheiris组上部发现了一种由分散的表皮组成的新的中观植物区系。该组合由许多不同形态的角质层组成，这些角质层均可分配给裸子植物。可以认定为corystospermalean类群角质层Dicroidium相当丰富，明显强于组装，而角质层分配给其他裸子植物是相当罕见的。迄今为止，所有三叠纪记录的Dicroidium这些表皮来自三叠纪古气候中35°S以上的古纬度地区，是该属的第一个记录。我们对该中观植物群的分析表明，在阿尼西斯晚期和世界其他地区一样，阿拉伯半岛陆地植物的裸子植物部分也存在。与产生各种针叶树花粉和其他裸子植物群花粉以及蕨类植物和苔藓植物丰富孢子的古植物相比，中植物群目前缺乏源植被中存在的许多植物群。 。差异很可能可以通过同源偏差来解释。表皮和古生物学的数据一起证明，至少在某种程度上，区域性陆地生态系统已经恢复。