Cuticles of Mariopteris occidentalis are described from the Desmoinesian (Middle Pennsylvanian) of Oklahoma (USA). This species, like other mariopteroids, had a vine- to liana-like growth habit and climbed with specialized climber hooks. However, M. occidentalis is different from other mariopteroids in having small recurved hooks on the abaxial surfaces of the pinna axes. The diagnosis for M. occidentalis White 1899 is emended based on additional macroscopical observations and data on the epidermal anatomy; a lectotype is designated. M. occidentalis is compared with Pseudomariopteris cordato-ovata from the Stephanian and Autunian of Europe and North America, a taxon which is considered very similar and may be related. Although the two taxa indeed display similarities, significant evidence for a closer relationship could not be found. Finally, some features, e.g. marginal water pits and the stomatal structure, are considered with regard to their palaeoautecological significance where they are interpreted as adaptations to special physiological requirements of a vine- to liana-like life form.

中文翻译：

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Mariopteris occidentalis White nov. 的角质层。订正。来自俄克拉荷马州中部宾夕法尼亚州（美国），以及一种用于悬翅类蕨类植物的新型登山钩

来自俄克拉荷马州（美国）的 Desmoinesian（宾夕法尼亚中部）描述了 Mariopteris occidentalis 的角质层。这个物种，像其他的旋翼类动物一样，有类似藤本植物的生长习性，并用专门的攀爬钩攀爬。然而，M. occidentalis 在耳廓轴的背面有小的反曲钩子，这与其他的悬翅类动物不同。M. occidentalis White 1899 的诊断基于额外的宏观观察和表皮解剖数据；指定选型。M. occidentalis 与来自欧洲和北美的 Stephanian 和 Autunian 的 Pseudomariopteris cordato-ovata 进行了比较，后者被认为非常相似并且可能相关。尽管这两个分类群确实显示出相似之处，但无法找到更密切关系的重要证据。最后，