Leaves of two new plants are reconstructed from their isolated leaflets collected from the Oligocene Los Ahuehuetes locality near Tepexi de Rodríguez in Puebla, Mexico. The leaves of Pseudosmodingium mirandae Ramírez‐Garduño et al. are compound imparipinnate with leaflets of variable morphology. The leaflets of five leaf morphotypes vary from narrow elliptic to lanceolate or lorate; they are symmetrical to slightly asymmetrical, with acute to attenuate apex, acute to cuneate base, and entire to serrate margin. Venation is simple pinnate craspedodromous, with secondary veins slightly curved near their base; secondary veins may dichotomize near the margin to become tertiary veins, and intersecondary veins are small and oblique to the secondary veins. A small number of leaflets assigned to Pseudosmodingium terrazasiae Ramírez‐Garduño et al. are distinguished from P. mirandae by the leaflet shape, length:width ratio, base shape, and apex angle. Morphological comparison of the fossil leaves with leaves of extant species of Anacardiaceae based on numerical analyses indicates a close similarity between P. mirandae and Pseudosmodingium multifolium Rose, while P. terrazasiae is more similar to Pseudosmodingium perniciosum (HBK) Engl. The presence of fossil species with extant relatives that are endemic to Mexico, along with previous reports, indicates that by the Oligocene, some lineages were already in place, although today they form part of the more xeric communities in southern North America.

中文翻译：

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墨西哥普埃布拉渐新世地层中两个新种紫菀（漆树科）叶片的重建

从墨西哥普埃布拉州 Tepexi de Rodríguez 附近渐新世 Los Ahuehuetes 地区收集的分离传单中，重建了两种新植物的叶子。Pseudosmodingium mirandae Ramírez-Garduño 等人的叶子。是具有可变形态的小叶的复合imparipinnate。五种叶片形态型的小叶从窄椭圆形到披针形或菱形；它们对称至略不对称，先端锐尖至渐细，基部锐尖至楔形，边缘全缘至锯齿状。脉是简单的羽状短脉，次脉在其基部附近略微弯曲；二级静脉在边缘附近可二分成为三级静脉，二级间静脉小且倾斜于二级静脉。分配给 Pseudosmodingium terrazasiae Ramírez-Garduño 等人的少量传单。与 P. mirandae 的区别在于小叶形状、长宽比、基部形状和顶角。基于数值分析的漆树科现存物种叶片化石叶的形态学比较表明 P. mirandae 和 Pseudosmodingium multifolium Rose 之间有密切的相似性，而 P. terrazasiae 更类似于 Pseudosmodingium perniciosum (HBK) Engl。具有墨西哥特有的现存亲属的化石物种的存在以及先前的报告表明，到渐新世，一些谱系已经存在，尽管今天它们构成了北美南部更加干旱的群落的一部分。基于数值分析的漆树科现存物种叶片化石叶的形态学比较表明 P. mirandae 和 Pseudosmodingium multifolium Rose 之间有密切的相似性，而 P. terrazasiae 更类似于 Pseudosmodingium perniciosum (HBK) Engl。具有墨西哥特有的现存亲属的化石物种的存在以及先前的报告表明，到渐新世，一些谱系已经存在，尽管今天它们构成了北美南部更加干旱的群落的一部分。基于数值分析的漆树科现存物种叶片化石叶的形态学比较表明 P. mirandae 和 Pseudosmodingium multifolium Rose 之间有密切的相似性，而 P. terrazasiae 更类似于 Pseudosmodingium perniciosum (HBK) Engl。具有墨西哥特有的现存亲属的化石物种的存在以及先前的报告表明，到渐新世，一些谱系已经存在，尽管今天它们构成了北美南部更加干旱的群落的一部分。