Periophthalmus barbarus Linnaeus, 1766 has many adaptations for amphibious life as a consequence of tidal zone occupation. One of them is the ability to keep a little amount of water and air in mouth while on land or in hypoxic water, correlated with closing a gill lid for gas exchange improvement. It causes that mechanisms of olfactory organ ventilation described in other species of actinopterygians (compression of accessory nasal sac(s) by the skull and jaw elements while mouth and gill lid moving) are not in operation. There is a specific mechanism of olfactory organ ventilation independent on jaw and skull elements movements. Compression of accessory nasal sacs is possible by a0 contraction and it is a movement effect on bones combined by ligaments. This process can be observed on P. barbarus as lifting the rostral part of the head.

中文翻译：

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眼周鱼（Gobiidae、Oxudercinae）嗅觉器官通气机制


由于占领潮汐带，Periophamus barbarus Linnaeus, 1766 具有许多适应两栖生活的能力。其中之一是在陆地上或缺氧水中时能够在口中保留少量的水和空气，这与关闭鳃盖以改善气体交换有关。它导致其他放线类物种中描述的嗅觉器官通气机制（当嘴和鳃盖移动时，头骨和下颌元件压缩副鼻囊）无法运行。嗅觉器官通气有一种独立于下巴和颅骨元件运动的特定机制。通过a0收缩可以压缩副鼻囊，这是对韧带结合的骨骼的运动效果。这个过程可以在 P. barbarus 上观察到，头部的喙部抬起。