Many enzymes, such as laccases, are involved in the saprotrophic lifestyle of fungi and the effects of those may be linked to enhanced bioweathering on stone surfaces. To test this hypothesis, we studied the decomposition of kerogen-enriched lithologies, especially with black slate containing up to 20% of C_org. Indeed, a formation of ditches with attached hyphal material could be observed. To address enzymes involved, proteomics was performed and one group of enzymes, the multicopper oxidase family members of laccases, was specifically investigated. A role in bioweathering of rocks containing high contents of organic carbon in the form of kerogen could be shown using the basidiomycete Schizophyllum commune, a white rot fungus that has been used as a model organism to study the role of filamentous basidiomycete fungi in bioweathering of black slate.

许多酶（例如漆酶）都参与了腐生真菌的生活，这些酶的作用可能与石材表面的生物风化作用增强有关。为了验证该假设，我们研究了富含干酪根的岩性的分解，特别是使用含有高达20％_Corg的黑色板岩的分解。实际上，可以观察到带有附着菌丝材料的沟渠的形成。为了解决涉及的酶，进行了蛋白质组学研究，并专门研究了一组酶，即漆酶的多铜氧化酶家族成员。可以使用担子菌裂殖体（Schizophyllum commune）来显示出以干酪根形式包含高有机碳含量的岩石在生物风化中的作用。，一种白色腐烂真菌，已被用作模型生物来研究丝状担子菌真菌在黑色板岩生物风化中的作用。