Based on an unexpected result of obtaining molecular sequence data from tropical representatives of the genus Normandina, we revised the biological concept of the neotropical taxon Marchandiomphalina foliacea. The obtained data let us conclude that M. foliacea is not a basidiomycete, as originally proposed, but belongs in Verrucariaceae, in the genus Agonimia, including its perithecia which had been identified with the lichenicolous Norrlinia peltigericola. The ITS (and nuLSU) sequences previously obtained from M. foliacea, seemingly confirming its status as a basidiomycete, are from an unmanifested lichenicolous fungus, present also in numerous specimens of Normandina. ITS data suggest the presence of seven lineages that can be recognized at the species level, forming two clusters: one cluster of three lineages found in thalli of M. foliacea, and a second cluster of four lineages found in thalli of Normandina. This pattern is similar to what has recently been found in the basidiomycete genus Cyphobasidium occurring predominantly in Parmeliaceae lichens. We propose the combination of Omphalina foliacea into the genus Agonimia, as Agonimia foliacea (P.M. Jørg.) Lücking & Moncada, comb. nov., and place Marchandiomphalina in synonymy with Agonimia. To formally recognize the unnamed lichenicolous basidiomycete present in Agonimia and Normandina thalli, we take advantage of provision ICN Art. 40.5 in the Code and describe the unmanifested fungus as a new genus, with seven new species, even if no physical type specimens can be preserved (except for the corresponding host lichens which, however, do not show the features of the fungus): Lawreymyces Lücking & Moncada, gen. nov. (Type: L. palicei), with L. bogotensis Lücking & Moncada, sp. nov., L. columbiensis Lücking & Moncada, sp. nov., L. confusus Lücking & Moncada, sp. nov., L, foliaceae Lücking & Moncada, sp. nov., L. palicei Lücking & Moncada, sp. nov., L. pulchellae Lücking & Moncada, sp. nov., and L. spribillei Lücking & Moncada, sp. nov. This opens the door to the formal recognition of thousands of species of voucherless fungi detected through environmental sequencing techniques under the current Code.

中文翻译：

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将Marchandiomphalina分解为Agonimia（Verrucariaceae）和Lawreymyces gen。十一月 （Corticiaceae）：为基于类型序列的数千种无券真菌的正式识别开创先例

基于从诺曼底属热带代表中获得分子序列数据的出乎意料的结果，我们修改了新热带分类群叶蓝线蛾（Marchandiomphalina foliacea）的生物学概念。所获得的数据让我们得出这样的结论M. foliacea不是一个担子，原来建议的，而是属于在Verrucariaceae，属中的Agonimia，包括它的子囊壳，其已确定与lichenicolous Norrlinia peltigericola。的ITS（和nuLSU）序列以前从获得M. foliacea似乎证实它是担子菌的地位，来自未显示的地衣真菌，也存在于诺曼底纳的许多标本中。ITS数据表明，存在七个可以在物种水平上识别的谱系，形成了两个簇：一个在三叶草的塔利中发现的三个谱系簇，另一个在诺曼底纳的塔利中发现的四个谱系簇。这种模式类似于最近一直在属担子菌发现Cyphobasidium主要发生在Parmeliaceae地衣。我们建议的组合Omphalina foliacea到属Agonimia，如Agonimia foliacea（PMJørg。）Lücking和Moncada，梳子。十一月 和地方Marchandiomphalina与同义Agonimia。为了正式认识存在于Agonimia和Normandina thalli中的未命名的地衣基底孢菌，我们利用ICN Art条款。《守则》中的40.5并将未显示的真菌描述为具有7个新物种的新属，即使无法保存任何物理类型的标本（但相应的寄主地衣除外，但其并未表现出真菌的特征）：Lawreymyces勒金＆蒙卡达，根。十一月 （Type：L. palicei），与波哥大乳杆菌吕金和蒙卡达，sp。十一月 ，L. columbiensis勒金＆蒙卡达，SP。十一月 ，L. confususLücking＆Moncada，sp.。十一月 ，L，叶植物Lücking＆Moncada，sp。十一月 ，L. palicei勒金＆蒙卡达，SP。十一月 ，L. pulchellaeLücking＆Moncada，sp。十一月 和L. spribilleiLücking＆Moncada，sp。十一月 这为正式承认通过现有规范通过环境测序技术检测的数千种无代价券真菌敞开了大门。