A multigene analysis of a combined ITS-LSU-SSU-rpb2-tef1 sequence data matrix was applied to infer the phylogenetic position of the genus Thyridaria in the Pleosporales. The generic type of Thyridaria, T. broussonetiae (syn. T. incrustans), is situated in a clade currently named Roussoellaceae, which becomes a synonym of Thyridariaceae. However, Thyridaria rubronotata does not belong to this clade, but is here recognised as Cyclothyriella rubronotata in its own family Cyclothyriellaceae. The Thyridariaceae contain the genera Thyridaria, Roussoella, Roussoellopsis, Neoroussoella and the new genus Parathyridaria. Roussoella acaciae is combined in Thyridaria and Roussoella percutaenea in Parathyridaria. Ohleria modesta and an additional new thyridaria-like genus, Hobus, are found to represent isolated lineages with unresolved phylogenetic affinites within the Pleosporales. For Ohleria the new family Ohleriaceae is established. Melanomma fuscidulum belongs to Nigrograna, and three new species are described in this genus. A strain named Biatriospora marina clusters with Nigrograna. Based on the newly recognised species in Nigrograna, morphology and ecology do in no way correlate among these genera, therefore we erect the new family Nigrogranaceae for Nigrograna and recommend to discontinue the use of the family name Biatriosporaceae until fresh material of B. marina becomes available for sequencing.

应用 ITS-LSU-SSU-rpb2-tef1 组合序列数据矩阵的多基因分析来推断胸孢菌属的系统发育位置。Thyridaria的通用类型，T. broussonetiae (syn. T. incrustans )，位于目前名为Roussoellaceae的进化枝中，它成为Thyridariaceae的同义词。然而，Thyridaria rubronotata不属于这个进化枝，但在这里被确认为Cyclothyriella rubronotata在它自己的科Cyclothyriellaceae中。稗科包含 胸腺 属,鲁梭氏 菌,鲁梭 氏 菌,新 鲁 梭 菌属 和 新 属副胸腺属. Roussoella acaciae在Thyridaria中组合在一起，Roussoella percutaenea在Parathyridaria中。发现奥氏菌属和一个额外的新的类似胸腺属的属Hobus代表了在 Pleosporales 内具有未解决的系统发育亲缘物的孤立谱系。为Ohleria建立了新的Ohleriaceae科。Melanomma fuscidulum属于Nigrograna，并在该属中描述了三个新种。一种名为Biatriospora marina的菌株与Nigrograna聚集在一起。基于新发现的Nigrograna物种，形态学和生态学在这些属之间没有任何关联，因此我们为 Nigrograna 建立了新的Nigrogranaceae并建议停止使用Biatriosporaceae直到B. marina的新鲜材料可用用于测序。