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The meiotic chromosomes (arrows) cluster away from the nuclear envelope in wild-type oocytes, but are closelyassociated with the nuclear envelope of Nup62 mutant oocytes. Loss of Nup62 disrupts global chromatin organisation in somatic cell lines, which is rescued by co-depletion of Nup155. Model for the chromatin attachment state controlled by an internal regulatory circuit in the NPC.