* Corresponding authors

^a Key Lab for Industrial Biocatalysis, Ministry of Education, Department of Chemical Engineering, Tsinghua University, Beijing 100084, P. R. China
E-mail: junge@mail.tsinghua.edu.cn

^b Institute of Biomedical Health Technology and Engineering, Shenzhen Bay Laboratory, Shenzhen 518107, P. R. China

^c State Key Laboratory of Bioreactor Engineering, Shanghai Collaborative Innovation Center for Biomanufacturing, East China University of Science and Technology (ECUST), Shanghai 200237, P. R. China

^d Department of Chemistry, Stanford University, Stanford, California 94305-5080, USA
E-mail: zare@stanford.edu

^e Institute of Biopharmaceutical and Health Engineering, Tsinghua Shenzhen International Graduate School, Shenzhen 518055, P. R. China