Abstract
The Devonian and Early Carboniferous brachiopod assemblages of the order Rhynchonellida from Transcaucasia (Nakhchivan Autonomous Republic of Azerbaijan and Armenia) are described. The succession of dominating rhynchonellid superfamilies and ecological types has been identified. Rhynchonellids were numerous in the Devonian; in the Early and Middle Devonian, the representatives of the superfamily Uncinuloidea prevailed. Their relict species survived up to the beginning of Famennian. The free-lying and anchoring cavern-dwelling rhynchonellids dominated in the Early and Middle Devonian. This ecological subtype is recognized for the first time. The rhynchonellid taxonomic and ecological groups changed at the Frasnian–Famennian boundary but rhynchonellids remained a significant part of the brachiopod fauna. The variability of rhynchonellids decreased at the Famennian–Tournaisian boundary but the superfamily Rhynchotrematoidea and the rhynchonellid anchoring ecological type continued to dominate starting from the Frasnian. The role of Rhynchonellida in the Early Carboniferous brachiopod fauna is insignificant. The decrease of variability within some zones is probably connected with taphonomic conditions.
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REFERENCES
Bolshakova, L.N., Hecker, M.R., Goryunova, R.V., Dubatolov, V.N., Ivanovskii, A.B., Kosmynin, V.N., Luchinina, V.A., Sokolov, B.S., and Tesakov, Yu.I., Paleozoic coral reefs on the territory of Russia, Stratigr. Geol. Korrel., 1994, vol. 2, no. 1, pp. 46–54.
Grechishnikova, I.A. and Levitskii, E.S., The Famennian-Lower Carboniferous Reference Section Geran-Kalasi (Nakhichevan Autonomous Region, Azerbaijan), Stratigr. Geol. Correl., 2011, vol. 19, no. 1, pp. 21–43.
Grechishnikova, I.A., Biostratigraphy of the Devonian and Carboniferous of Transcaucasia (Brachiopod Zonal Scale), Paleontol. J., 2018, vol. 52, no. 8, pp. 832–862.
Ivanova, E.A., Conditions of existence, mode of life, and history of the development of some brachiopods from the Middle and Upper Carboniferous of the Moscow Depression, Tr. Paleontol. Inst. Akad Nauk SSSR, 1949, vol. 21, pp. 1–152.
Jackson, J.B.C., Goreau, T.F. and Hartman, W.D., Recent brachiopod – coralline sponge communites and their paleoecological significance, Science, 1971, vol. 173, pp. 623–625.
Logan, A. and Zibrowius, H., A new genus and species of rhynchonellids (Brachiopoda, recent) from submarine caves in the Mediterranean Sea, Publ. Staz. Zool. Napoli:Mar. Ecol., 1994, vol. 15, no. 1, pp. 77–88.
Logan, A., Ecological observations on the recent articulate brachiopod Argyrotheca bermudana Dall, from the Bermuda platform, Bull. Biol. Sci., 1975, vol. 25, pp. 186–204.
Makkaveeva, E.V., Naselenie nekotorykh banok Egeiskogo morya [Inhabitants of some banks in the Aegean Sea], Tr. Sevastopol’ Biol. St., 1963, vol. 16, pp. 211–214.
Mirieva, G.T., Zonal’noe raschlenenie otlozhenii verkhnego devona Yuzhnogo Zakavkaz’ya (Nakhchyvanskoi AR) po rinkhonellidam [Zonal subdivision of the Upper Devonian deposits in the Southern Transcaucasia (Nakhchivan AR) by rhynchonellids], The evolution of the organic world and biotic crises. Mat. LVI s. Pal. Soc. RAS (5th–9th April 2010, St. Petersburg), pp. 74–76.
Pakhnevich, A.V., Development of punctation in the shells of Late Devonian brachiopods of the order Rhynchonellida, in Tezisy konferentsii “Morfogenez v individual’nomi istoricheskom razvitii: ustoichivost’ i variabel’nost’”, 21–23 aprelya 2015 g. (Conference on the Morphogenesis in Individual and Historical Development: Stability and Variability, April 21–23, 2015), Moscow: Paleontol. Inst. Ross.Akad. Nauk, 2015, pp. 50–51.
Pakhnevich, A.V., Order Rhynchonellida Kuhn, 1949, Paleontol. J., 2018, vol. 52, no. 8, pp. 892–956.
Pakhnevich, A.V., Reasons of micromorphism in modern and fossil brachiopods, Paleontol. J., 2009, vol. 43, no. 11, pp. 1458–1468.
Ruggiero, E.T., Brachiopods of the Isca submarine cave: observations during ten years, 4th Intern. Brachiopod Congr., London, Great Britain, Vol.: The Millennium Brachiopod Congress. Abstracts. L.: Natur. Hist. Mus., 2000, p. 74.
Rzhonsnitskaya, M.A. and Mamedov, A.B., Devonian stage boundaries in the southern Transcaucasus, Cour. Forsch.-Inst. Senckenb., 2000, vol. 225, pp. 329–333.
Saito, M., Motchorova-Dekova, N. and Endo, K., Recent brachiopod fauna from the submarine caves of Okinawa, Japan, 4th Intern. Brachiopod Congr., London, Great Britain, Vol.: The Millennium Brachiopod Congress. Abstracts. London: Natur. Hist. Mus., 2000, pp. 77.
Simon, E. and Willems, G., Gwynia capsula (Jeffreys, 1859) and other recent brachiopods from submarine caves in Croatia, Bull. Inst. Roy. Sci. Natur. Belg. Biol., 1999, vol. 69, pp. 15–21.
Swedmark, B., The interstitial fauna of marine sand, Biol. Rev., 1964, vol. 39, pp. 1–42.
Zezina, O.N. and Pakhnevich, A.V., On finding of recent articulate brachiopods at underwater volcanic activity region of the Antarctic Peninsula, Dokl. Akad. Nauk SSSR, 2000, vol. 370, no. 3, pp. 397–398.
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Pakhnevich, A.V. Ecological and Taxonomical Succession of Rhynchonellids (Brachiopoda) in the Devonian and Early Carboniferous of Transcaucasia. Paleontol. J. 54, 584–593 (2020). https://doi.org/10.1134/S0031030120060064
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DOI: https://doi.org/10.1134/S0031030120060064