Japanese species of the genus Woldstedtius Carlson, 1979 are revised. Nine species are recorded from Japan, including two new species, W. alpicolasp. nov. and W. punctatussp. nov.Woldstedtius biguttatus (Gravenhorst, 1829) is newly recorded from Japan. Taxonomic status of W. flavolineatus kuroashii (Uchida, 1957) is changed from the subspecies of W. flavolineatus (Gravenhorst, 1829) to a separated

 Of the 12 specimens of Calathus-like sphodrine beetles presently known from Baltic and Rovno amber deposits, 11 specimens were investigated using light microscopy, micro-CT scanning, and 3D imaging techniques. For the first time, many significant diagnostic characters of the external morphology and male and female genitalia of Eocene Sphodrini were studied in detail. Based on these data, three fossil

 In spite of big data and new techniques, the phylogeny and timing of cockroaches remain in dispute. Apart from sequencing more species, an alternative way to improve the phylogenetic inference and time estimation is to improve the quality of data, calibrations and analytical procedure. This study emphasizes the completeness of data, the reliability of genes (judged via alignment ambiguity and substitution

A new fossil species of the extant archostematan genus Omma Newman, O. fortesp. nov., is reported from mid-Cretaceous amber from northern Myanmar. The extinct ommatid genus, Cionocups Kirejtshuk from the same deposit, is considered as a junior synonym of Omma, and C. manukyani is transferred to Omma, as O. manukyanicomb. nov. A key to species of Omma from Burmese amber is also provided.

 The species of the genus Phanuromyia Dodd, 1914, belonging to the subfamily Telenominae are egg parasitoids of planthoppers belonging to the families Issidae, Flatidae, Fulgoridae and Ricaniidae (Hemiptera: Fulguroidea). So far eleven species of Phanuromyia are known from India. Fifteen new species of Phanuromyia Dodd are described here: P. chalukyasp. nov., P. cherasp. nov., P. cholasp. nov., P

The relictual archostematan beetle family Ommatidae attained high diversity during the Mesozoic. Despite their once high taxonomic diversity and morphological disparity, many Mesozoic ommatid taxa remain poorly understood, partly due to limited preservation. Here we report an exceptionally well-preserved fossil, which we describe as a new ommatid genus and species, Limnomma daohugouensegen. et sp.

Japanese species of the genus Xanthopimpla Saussure, 1892 are revised. A total of 15 species are recorded from Japan, including three new species, X. nipponensissp. nov. X. sylvicolasp. nov. and X. yoshimuraisp. nov. and three species, X. honorata (Cameron, 1899), X. minuta Cameron, 1905 and X. trias Townes & Chiu, 1970, newly recorded from Japan. Host, habitat, overwintering and distribution patterns

The Iranian species of the genus Schoenlandella Cameron, 1905 are revised based on morphological characters. The current work provides a key to species of Iranian Schoenlandella and species descriptions of two new species: S. angustigena Kang sp. nov. and S. latigena Kang sp. nov. This work results in increasing the number of Iranian Schoenlandella species from one to three species.

Ornatoraphidia flavilabris (Costa, 1851) is one of 15 snakefly species occurring in southern parts of Central Europe. It is a polycentric Mediterranean faunal element with refugia in the Apennine Peninsula and the Balkan Peninsula. Two phylogeographic questions are dealt with in this paper: (1) Is it possible to differentiate, morphologically or genetically, the Balkanic populations from the Italian

The Pyrenees are inhabited by scattered populations of earwigs of the genus Chelidura Latreille, 1825. There is some controversy about the specific assignment of these populations: while most authors assign them to C. pyrenaica (Gené, 1832), other consider that C. aptera (Mégerlé, 1825) is also present in the Pyrenees. The main objective of this work was to revise the identity and synonyms of Pyrenean

The myrmicine genus Rotastruma is a rarely collected arboreal ant group and only two species, R. recava Bolton and R. stenoceps Bolton, are known from the Oriental Region. A new species, R. epispinasp. nov., is described from Cambodia based on the worker and queen castes. The new species is distinguished from R. recava and R. stenoceps by having the propodeal spine directed upwards and elongated peduncle

Balticeler kerneggerigen. nov., sp. nov., is described based on six fossil specimens preserved in Eocene Baltic amber and imaged using light microscopy and X-ray micro-computed tomography. Based on certain characters observed in the fossil species it is considered a “middle grade” Carabidae, outside of the large family Harpalinae (as it possesses a scrobal seta, the lack of which is a synapomorphy

Four new species of Agraphydrus Régimbart are described from China: A. pseudonigersp. nov. from Shangyou County, Jiangxi Province, A. komarekisp. nov. from Shangchuan island, Taishan County, Guangdong Province, A. sabulosussp. nov. from Fengkai, Guangdong Province, A. dapengensissp. nov. from Dapeng peninsula, Shenzhen, Guangdong Province. Diagnosis and illustration of the new species are provided

The first instar nymphs of two peltoperlid stoneflies, i.e., Microperla brevicauda Kawai, 1958 of Microperlinae and Yoraperla uenoi (Kohno, 1946) of Peltoperlinae, were examined and described. Additionally, the phylogeny and groundplan of the first instar nymphs of Peltoperlidae and Plecoptera were considered. The first instar nymphs of M. brevicauda have a slender body with a prognathous head of typical

A new monotypic genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae) from New Georgia Island (Solomon Islands), Kazukurugen. nov., is described for K. zingiberissp. nov. (type species). Habitus, female, external and internal genital structures of the new species are described and illustrated.

During several excursions to the Chin Hills of Myanmar from 2001 to 2015, rich collections of caddisflies were made which form the basis of the present taxonomic and biogeographic study. A total of 106 species were identified including seven new species. They are described as Arctopsyche subflavasp. nov., Hydromanicus abdominalissp. nov., Cheumatopsyche janosolahisp. nov., Lepidostoma subpanaitossp

Most species of the Colletes succinctus group sensu Noskiewicz, 1936 are taxonomically uncertain. This study has chosen an integrative approach, including pollen analysis, morphology, male genitalia, morphometry, cuticle sculpture and DNA-barcoding (CO1) to investigate the five species that were reported from Austria. It includes a detailed analysis of the male genitalia and the first description of

Species of Bibionidae from Baltic amber are reevaluated based on newly discovered material, and a key to the species is given. Bibio succineussp. nov. is described based on one male specimen, this is the first Bibio named from Baltic amber. The males of Hesperinus electrus Skartveit, 2009 and Penthetria montanaregis Skartveit, 2009 are redescribed. A single, autoclave treated specimen of Penthetria

The genera Cestromoecha and Poreuomena of the tribe Poreuomenini in Phaneropterinae are revised and new generic characters are given for both genera, and six new species are described in Poreuomena. The newly described species are P. biaculeatasp. nov., P. ealasp. nov., P. gracilicercatasp. nov., P. ivorianasp. nov., P. matthaeisp. nov., and P. tshuapasp. nov. Based on characters defining the two genera

The world species of the microgastrine genus Paroplitis (Hymenoptera: Braconidae) are revised. Three new species are described, P. horticola Fujie & Fernandez-Triana, sp. nov. and P. japonicus Fujie & Fernandez-Triana, sp. nov. from Japan and P. kakhetiensis Fujie, Japoshvili & Fernandez-Triana, sp. nov. from Georgia. P. vietnamensis van Achterberg & Fernandez-Triana, 2013 is re-described, based on

The sting apparatus of aculeate Hymenoptera is derived from the ovipositor and is their most prominent apomorphy. In contrast to the frequently analysed sclerites of the sting apparatus, the associated musculature has largely been neglected. In this study, we use micro-computed tomography to present a detailed description of the musculature of the sting apparatus of Ampulex compressa (Ampulicidae)

Investigations of type material and new material from the Philippines (Luzon) revealed that Baetis luzonensis Müller-Liebenau, 1982 and B. realonae Müller-Liebenau, 1982 do not belong to Baetis Leach, 1815. A new genus, Philibaetis gen. nov., is described to accommodate both species and both are re-described based on larvae. The new genus is characterised by having a rectangular labrum with a submarginal

The species of three subgenera of the genus Bracon Fabricius (Hymenoptera, Braconidae, Braconinae), i.e. Glabrobracon Fahringer, 1927; Lucobracon Fahringer, 1927; and Uncobracon Papp, 1996 from China are revised and 31 species are recognised, including 11 new species, i.e. Bracon (Glabrobracon) indistinctus sp. nov., B. (G.) leptotes sp. nov., B. (G.) longistriatus sp. nov., B. (G.) megaventris sp

The ground beetle genus Agonum Bonelli, 1810 is a large genus of the tribe Platynini with many species that show high amounts of intraspecific variations, making a correct identification challenging. As part of the German Barcode of Life initiative, this publication provides a comprehensive DNA barcode library for species of Agonum that are reported for Germany. In total, DNA barcodes from 258 beetles

Using high-resolution stereomicroscopy and exploratory data analyses, a taxonomic revision of the cryptic species close to Plagiolepis schmitzii Forel, 1895, called Pl. schmitzii group, was conducted. Morphology was numerically recorded under highly standardised conditions considering absolute size and 16 shape, pubescence and surface characters. A key to the non-parasitic Westpalaearctic species of

Bayesian and parsimony phylogenetic analyses of combined and partitioned datasets of molecular (partial sequences of 28S, wg, COI, and CAD) and morphological (51 characters of adults) data for exemplar taxa of five outgroup and 76 ingroup abacetine carabids resulted in a monophyletic Loxandrina Erwin & Sims, 1984 that is split into Australian and American clades. The genus Loxandrus LeConte, 1853 as

Male and female of Nusalala peruanasp. nov., a flightless hemerobiid from the Andes mountain range of northern Peru, at a height of almost 4000 m, are described, figured and documented as the first record of a brachypterous, flightless species of Nusalala Navás, 1913, from this country. The other two congeneric, brachypterous species are from high altitudes in Colombia and Costa Rica and have been

Two tenthredinid sawfly species are newly recorded in Germany: Pristiphora krausi (Lacourt, 2006) and P. melagonia sp. nov. The latter is also recorded from Greece and Spain. These species belong to the Pristiphora depressa group, as confirmed by genetic data. Additional data are presented for seven other tenthredinid species which have only rarely been recorded in Germany and for which previously-published

A total of 10 species of Enicospilus (Hymenoptera, Ichneumonidae, Ophioninae) have previously been reported from Nepal. Six new species are described here (E. alleni Shimizu sp. nov., E. kakanicus Shimizu sp. nov., E. nepalensis Shimizu sp. nov., E. nikami Shimizu sp. nov., E. phulchokiensis Shimizu sp. nov., and E. tangi Shimizu sp. nov.), and 10 are newly recorded (E. ashbyi Ashmead, 1904, E. bifasciatus

Conflicting hypotheses about the relationships among the major lineages of aculeate Hymenoptera clearly show the necessity of detailed comparative morphological studies. Using micro-computed tomography and 3D reconstructions, the skeletal musculature of the mesoand metathorax and the first and second abdominal segment in Apoidea are described. Females of Sceliphron destillatorium, Sphex (Fernaldina)

Himalcercyon Hebauer, 2002 stat. nov. is elevated to genus rank based on the unique form of its mesoventral elevation. The genus is reviewed, redescribed, and illustrated in detail. Two species are recognized: Himalcercyon mirus (Hebauer, 2002) comb. nov. from Nepal and H. franzi sp. nov. from Chinese Himalaya (Xizang Autonomous Region). Both species are illustrated and diagnosed. An updated key to

Tomares hairstreaks comprise about 10 species distributed from Europe and North Africa to Central Asia. The taxonomy of the genus is hampered by the absence of diagnostic characters by which specimens can be unambiguously assigned to species. Our investigation of morphology and DNA barcode variations within and between Tomares species shows that while well-defined species (T. ballus, T. mauritanicus

Very few sawflies using Gentianaceae as larval host plants have been recorded. We identified larvae collected in Austria on Gentiana asclepiadea L. as Tenthredo atra Linnaeus, 1758 and T. propinqua Klug, 1817. If its current taxonomic circumscription as a single species is accepted, T. atra is a highly polyphagous species, whereas T. propinqua may be more specialised: Gentiana asclepiadea is its first

The leafhopper genus Opsius Fieber, 1866 is revised for the Kingdom of Saudi Arabia. Seven species are treated, including three that previously were reported by Dlabola (1979), O. pallasi (Lethierry, 1874), O. tigripes (Lethierry, 1876), and O. versicolor (Distant, 1908). Opsius heydeni (Lethierry, 1876), O. richteri Dlabola 1960, and O. scutellaris (Lethierry, 1874) are reported for the first time

Leafand fruit-feeding chrysomelids (Coleoptera) on bananas and plantains (Musaceae, Zingiberales) cause major losses to banana growers in the northern and northeastern regions of India, Bangladesh, and other parts of Southeast Asia. The species composition of these beetles has not been studied so far in India and wrong names or wrong name combinations in the literature have caused confusion. Most particularly

Tropopterus Solier, 1849, precinctive to southern South America, is taxonomically revised. Six new species are described: T. peckorum sp. nov., T. robustus sp. nov., T. canaliculus sp. nov., T. trisinuatus sp. nov., T. minimucro sp. nov., and T. fieldianus sp. nov. Merizodus catapileanus Jeannel, 1962, is synonymized with T. montagnei Solier, 1849. Lectotypes are designated for T. montagnei, T. giraudyi

Here we elucidate and justify a DNA barcode approach to insect species description that can be applied to name tens of thousands of species of Ichneumonoidea and many other species-rich taxa. Each description consists of a lateral habitus image of the specimen, a COI barcode diagnosis, and the holotype specimen information required by the International Code of Zoological Nomenclature. We believe this

The larval morphology of the water scavenger beetle Psalitrus yamatensis Hoshina & Satô, 2005 is described based on a specimen collected from Fukuoka Prefecture, Kyushu, Japan. This is the first description of the larval morphology of the hydrophilid genus Psalitrus d’Orchymont, 1919, as well as the first description of larval chaetotaxy of the tribe Omicrini. Species-level identification of the larva

Thirty-nine species of sawfly (Symphyta) are recorded for the first time in Bulgaria. Most of these were collected during early spring of 2018, in the south-east of the country (Burgas and Varna Provinces). Empria aridicola Macek & Prous, sp. nov. is described as new to science from specimens collected in several central, east and south European countries. Lectotypes are designated for Poecilosoma

A new and a previously known species of the genus Zhenia Q. Zhang, 2016 (Eremochaetidae) are illustrated and described based on two males and a female in amber: Zhenia burmensis sp. nov. and Z. xiai Q. Zhang, 2016. The male Z. xiai is the first male of this species recorded. The relationships of Archisargoidea (including Eremochaetidae, Zhenia) are reassessed based on male genitalia. The superfamily

The genus Notiophilus Duméril, 1806 is a distinctive taxon of small, diurnal and morphologically similar beetles exhibiting large eyes and widened second elytral intervals. In this study we analysed the effectiveness of DNA barcodes to discriminate 67 specimens that represent 8 species of Notiophilus from Central Europe. Interspecific K2P distances below 2.2% were found for N. biguttatus (Fabricius

A study of numeric morphology-based alpha-taxonomy (NUMOBAT) considering the species Formica exsecta Nylander, 1846 and F. fennica Seifert, 2000 was performed in 166 nest samples with 485 worker individuals originating from 117 localities of the Palaearctic west of 59°E. The presence of intraspecific pilosity dimorphism is shown for F. exsecta. The setae-reduced phenotype, termed the Rubens morph,

Type material of thick-headed flies (Diptera, Conopidae) in the collection of the Museum für Naturkunde Berlin (ZMHB) is documented. The entire collection holds primary type material (i.e. holotypes, lectotypes, syntypes) of 73 species. Five Conops species with previously unknown subgenus belong to the subgenus Asiconops: C. frontosus Kröber, 1916; C. indicus Kröber, 1916; C. maculiventris Kröber,

Thirteen species of chewing lice in the Brueelia-complex are redescribed and illustrated. They are: Brueelia blagovescenskyi Balát, 1955, ex Emberiza schoeniclus (Linnaeus, 1758); B. breueri Balát, 1955, ex Chloris chloris (Linnaeus, 1758); B. conocephala (Blagoveshchensky, 1940) ex Sitta europaea (Linnaeus, 1758); B. ferianci Balát, 1955, ex Anthus trivialis (Linnaeus, 1758); B. glizi Balát, 1955

Almost 200 years of continuous and systematic research in subterranean habitats of the Dinaric Karst and adjoining areas have resulted in the discovery of more than 400 specialized subterranean beetles. Among these, a special place belongs to the morphologically well distinguished and elusive, so called aphaenopsoid trechine beetles, which are characterized by a prolonged head, pronotum and appendages

Based on cladistic analyses recently conducted by Sharkey and Chapman, the genus Zosteragathis Sharkey is revised. Twenty-two species are reported from Thailand, three previously described species, Z. samensis, Z. contrasta and Z. nuichuaensis, and 19 new species, i.e., Z. chaiyaphumensis, Z. eukos, Z. hinensis, Z. hongensis, Z. inthanonensis, Z. krachanensis, Z. lampangensis, Z. lampooensis, Z. luangensis

The Australian fauna of Mecyclothorax Sharp (Coleoptera: Carabidae: Moriomorphini) is reviewed, with special focus on species assigned to the monophyletic subgenus Eucyclothorax Liebherr: M. isolatus, sp. n. from Western Australia, M. moorei Baehr, M. punctatus (Sloane), M. curtus (Sloane), M. blackburni (Sloane); M. eyrensis (Blackburn); M. peryphoides (Blackburn); M. darlingtoni, sp. n. from Queensland;

The authors revised the genus Morgenia Karsch, 1890 which now consists of eight species, of which three are here newly described (Morgenia plurimaculata Massa & Moulin, sp. n., M. angustipinnata Massa, sp. n., and M. lehmannorum Heller & Massa, sp. n.). Six of the eight species occur in the Tri National Sangha (TNS) comprising Dzanga-Sangha Special Reserve and Dzanga Ndoki National Park (Central African

Twenty eight species of the genus Quedius from Middle Asia comprising Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan, are revised. Quedius altaicus Korge, 1962, Q. capitalis Eppelsheim, 1892, Q. fusicornis Luze, 1904, Q. solskyi Luze, 1904 and Q. cohaesus Eppelsheim, 1888 are redescribed. The following new synonymies are established: Q. solskyi Luze, 1904 = Q. asiaticus Bernhauer,

An improved diagnosis of the genus Tricerophora is provided, and its position within Gelechiidae is briefly discussed. A new generic synonym is established: Leucophylla Janse, 1960, syn. n. of Tricerophora Janse, 1958. Six new species are described: T. pundamilia sp. n. (RSA), T. rukinga sp. n. (Kenya), T. nigrinervis sp. n. (RSA, Namibia), T. brumale sp. n. (Namibia), T. acutivalva sp. n. (Iran),

The hydrophilid genus Cycreon Orchymont, 1919, previously known from two historical specimens only, is reviewed based on the numerous material collected recently from the inflorescences of various Araceae species in the Malay Peninsula and Borneo. Four species are recognized in the genus: C. sculpturatus Orchymont, 1919 from Sumatra, C. armandi Shatrovskiy, 2017 from Singapore, C. adolescens sp. n

The genus Vansoniella gen. n. is established to accommodate the species V. chirindensis sp. n., collected in Zimbabwe by Van Son in 1937. The new species differs externally from other African taxa by translucent foreand hindwings in the male sex. The wing venation is highly derived and the male genitalia are also structurally different from other genera. The genus occupies an isolated position within

Social wasps perform a variety of tasks with their mouthparts. Female workers use them to feed on carbohydrate-rich fluids, to build nests by collecting wood fibers and forming paper, to hunt and manipulate insect prey for feeding larvae as well as for brood care. Since male wasps neither feed on insects nor participate in nest building, sex-specific differences in mouthpart morphology are expected

The 15 species of Mecyclothorax Sharp precinctive to New Caledonia are revised and shown by cladistic analysis to comprise a monophyletic lineage, here treated as subgenus Phacothorax Jeannel. The New Caledonian species of subgenus Phacothorax include Mecyclothorax fleutiauxi (Jeannel), M. najtae Deuve, and 13 newly described species: M. jeanneli sp. n., M. laterobustus sp. n., M. laterorectus sp.

This year we look back at 160 years of entomological research published in the DEZ. Believe it or not, our journal is the third oldest of all still existing entomological periodicals worldwide! A concatenation of favourable circumstances? At first glance, the first decades were rather tough ones, involving personal controversies, splitting of the society behind the journal and the journal itself, and

The revised checklist of the subtribe Poliina Hampson, 1902 is presented; one new genus, Multisigna gen. n., three new subgenera (Atropolia, Leuconephropolia and Protopolia subgen. n.) and a new species (Polia (Atropolia) posterodiluta sp. n.) are described. The taxonomic position of the recently described subgenus Metallopolia is discussed. The subtribe Pachetrina Beck, 1996 is synonymised with Poliina;

The Mediterranean snakefly Raphidia mediterranea H. Aspöck, U. Aspöck & Rausch, 1977 – known from many parts of the Balkan Peninsula, several Aegean islands, southern parts of Italy, northwest of Anatolia and a few localities in Eastern Europe, yet not recorded in Central Europe – was surprisingly found with an astoundingly high population density on bushes in the yard of an old farmhouse at a comparatively

This monographic review of the Nevrorthidae Nakahara, 1915, covers all 19 validly described, extant species worldwide that belong to one of the smallest families of the order Neuroptera. The family embraces four genera: Nevrorthus Costa, 1863 (with five species occurring in the Mediterranean region), Austroneurorthus Nakahara, 1958 (with two species restricted to eastern Australia), Nipponeurorthus

A new species of a new subgenus and a similar known species referred to the genus Flagellisargus J Zhang, 2012 are described and illustrated based on a male and a female impression fossils of these flies: Flagellisargus (Changbingisargus) parvus subgen. et sp. n. and Flagellisargus (Flagellisargus) cf. sinicus J Zhang, 2012. The latter taxon is the first record of a female Flagellisargus. Recently

The cuckoo wasp genus Pseudochrysis Semenov, 1891 is currently treated by several authors as a junior subjective synonym of Euchroeus Latreille, 1809, due to a type species designation by O. W. Richards in 1935. In the original description of the genus Pseudochrysis, Semenov (1891) distinguished two subordinated taxa within the genus Pseudochrysis: the subgenus Pseudochrysis and the subgenus Spintharis