The uppermost Pennsylvanian Pony Creek Shale Lagerstätte (Pony Creek Shale Member, Wood Siding Formation, Wabaunsee Group) near Maple Hill, Wabaunsee County, Kansas, contains exceptionally preserved horseshoe crab (Xiphosura) fossils. Associated with these fossils is an ichnofauna that includes trace fossils assumed to be produced by xiphosurids, as well as coprolites and a few burrows. Of the trace

(2020). One legacy: ONE ICHNOLOGY! Richard Granville Bromley (1939–2018) Ichnos: Vol. 27, Richard G. Bromley Memorial Issue, pp. 249-257.

Bromlichnus bromleyi n.igen. n.isp., a compound trace fossil consisting of two segments representing dwelling and feeding, is described. Owing to a concentric spreite in a bow-shaped burrow, B. bromleyi is interpreted as made by holothurians. In most cases, it is connected to a repichnion. The new ichnotaxon derives from the narrow Pfalzpaint Subbasin of the Solnhofen archipelago (Upper Jurassic, SE

Many aspects of the palaeontology of the Isle of Wight are well known, but less so its ichnology. A turtle bone, Emys? sp., from the Lower Oligocene (Rupelian) Hamstead Member, Bouldnor Formation, exposed on the north coast of the island, preserves an unusual surface etching. Dictyoporus nodosus Mägdefrau is recorded from the Paleogene for the first time; hitherto, in northern Europe, it was only known

The design of a relational database for ichnological data is presented to illustrate and address deficiencies in present-day palaeontological databases. Currently, palaeontology databases apply concepts and terminology derived from the study of body fossils to trace fossil records. We suggest that fundamental differences between body and trace fossils make this practice inappropriate. These differences

Disconnected aggregates of allochthonous fossil Nypa fruits occur in the Ammonite Hill Member of the Dakhla Formation (Maastrichtian), exposed at the Bir Abu Minqar section, south Western Desert of Egypt. In a pale brown, calcareous siltstone and/or fine-grained sandstone layer, the fossil fruits found are densely infested by the borings Teredolites clavatus and Apectoichnus longissimus. In the specimens

The Mukdadiya Formation (Late Miocene–Pliocene) consists of alternations of red to brown sandstone and claystone layers in the Zagros Mountains Belt, northeastern Iraq. Two tracksites preserving bird and mammal tracks were recorded from the base of the formation in the Chamchamal area of the Kurdistan region in northeast Iraq. Avian tracks are large footprints with an average length of 25 cm that belong

Ichnofossils from the type locality of the Famennian are here described and figured for the first time. The siliciclastic supratidal alluvial to proximal subtidal deposits of S Belgium yield a diverse ichnofauna composed of Arenicolites ispp. (two morphotypes), Chondrites intricatus, Diplocraterion isp., Dolopichnus gulosus, Gordia marina, Helminthopsis isp., Lockeia cordata, L. siliquaria, Palaeophycus

A proximal distributary channel deposit in the Turonian Ferron Sandstone of central Utah contains a trace-fossil assemblage comprised of turtle tracks (Chelonipus), vertebrate swim trace fossils (Characichnos), repichnial traces of mollusks (Archaeonassa), Rhizocorallium, and Teredolites. Mayflies are abundant producers of horizontal, spreiten, U-shaped burrows in modern freshwater channels, but rarely

A new high-latitude record of the trace fossil Macaronichnus segregatis degiberti is documented from the lower Miocene Punta Basílica beds, Tierra del Fuego, which were previously considered to be deep-marine channel and levee deposits. Based on a combined ichnological and sedimentological study, however, these beds are herein re-interpreted as shallow-marine, delta-front clinoform deposits. Macaronichnus-bearing

(2020). Correction. Ichnos: Vol. 27, No. 1, pp. 80-80.

The association of vertebrate remains and invertebrate traces, although less studied than other bioerosion traces, provides important paleoecological information. This report describes Cubiculum ornatum Roberts, Rogers, and Foreman 2007 Roberts, E., R. Rogers, and B. Foreman. 2007. Continental insect borings in dinosaur bone: examples from the Late Cretaceous of Madagascar and Utah. Journal of Paleontology

The large holasteroid echinoid Hemipneustes striatoradiatus (Leske) was exploited by diverse invertebrate encrusters and borers during the Maastrichtian, both pre- and post-mortem. In life, the specimen described herein was perforated by multiple Oichnus simplex Bromley borings close to the apical system. Each engendered a growth reaction from the echinoid, a mound-like swelling on the external surface

Jurassic deposits of shallow to marginal marine (delta) environments are widely reported from different continents of the world. This study shows inter-relationship of the animal-sediment behaviours in shallow and marginal marine conditions, suggesting an interpretation of the possible ichnodisparity. The Jurassic succession exposed at Washtawa Dome and Adhoi Anticline of Wagad highland, Kachchh comprises

Integrated ichnology, sedimentology and sequence stratigraphy of the Lower Quartzite Member to the Arkosic Sandstone Member of the Koti Dhaman Formation (Cambrian Series 2, Stage 4), Tal Group, Nigali Dhar Syncline, Lesser Himalayan lithotectonic zone are presented. Trilobite traces of Gondwanan affinity i.e., Cruziana salomonis, Cruziana fasciculata, Rusophycus dispar and Rusophycus burjensis are

This study presents an ichnotaxonomical assessment of bird footprints from a coastal plain setting in the Eocene lower to middle Aspelintoppen Formation in Brongniartfjellet and Storvola, Svalbard. These footprints are unique evidence for Paleogene birds from Svalbard and the second evidence of Paleogene avifauna of the Arctic (along with scarce fossil remains from Ellesmere Island). The analyzed footprints

Microcoprolites of Palaxius salataensis are non-skeletal carbonate grains produced by callianassid shrimps. This ichnospecies is recognized by four longitudinal canals arranged around a bilateral plane of symmetry, with crescent outline in cross-section. Palaxius salataensis has been previously reported from other countries and in strata from the late Paleozoic to the Mesozoic. In this study, we describe

In his short joint ventures across the world of vertebrate tracks, Richard Bromley recognized the aeolian sands as unsuitable soft substrates for their preservation. Only after his work in the Balearic Islands, a more systematic study of coastal aeolianites worldwide revealed that these depositional systems could preserve a highly important record of behavioural trace fossils for the evolution of vertebrates

Bowl-shaped structures are for the first time reported from the Greek Island of Rhodes. They occur in Pleistocene deposits of the Cape Arkhangelos Formation in the Rhodes Synthem. The regularity of their three-dimensional appearance is the argument for a biological origin. This gives the reason to assign the structures to Piscichnus waitemata for which we issue a formal diagnosis. They were formed

Many tracemakers use different materials to line their burrows. Koptichnus rasmussenae n. igen. n. isp. is lined with cuboid fragments of siliceous sponges, interpreted as evidence of harvesting and trimming material to reinforce the burrow wall. The act of trimming, as evidenced in the polyhedral faces, is considered to be behaviourally significant. The tracemaker was evidently a lobster-like crustacean

Scolicia is one of the most conspicuous trace fossils in lower shoreface deposits of the Miocene Chenque Formation of Patagonia, Argentina. This ichnotaxon consists of horizontal, sinuous or meandering trace fossils with a laminated backfill and two parallel strings located at the base. Abundant body fossils attributed to Brisaster iheringi occur in close association to these trace fossils. The echinoids

A new ichnospecies, Linichnus bromleyi, is described on bone substrate as the result of a very likely predator/scavenger interaction. L. bromleyi consists of a single groove with a non-serrated edge. This new ichnotaxon is compared with L. serratus which was defined as a single elongate serrate-edged groove. L. bromleyi has been identified over the surface of bones of marine mammals (in particular

An insect boring of unique shape is described from a lignitic layer within the Early Cretaceous (Berriasian) Skyttegård Member of the Rabekke Formation on Bornholm. Morphologically it cannot be compared to any modern or fossil wood borings, although some structures are reminiscent of Scolytidae, Platypodidae and Lymexylonidae. Most probably, however, the tracemaker was a female fungus-farming beetle

Peculiar meniscate burrows with three sediment cords occur in early to middle Miocene tidal-flat deposits of southwestern Japan. Two of the cords are situated at the bottom and the other is at its center. Detailed observations of the burrow structures and comparative neoichnological studies of modern spatangoid burrows in a tidal flat revealed that the former two were true drainage tubes and the latter

Image processing techniques, including the Analyze Particles tool offered by Fiji software and the Intensity Profile by ICY (IP-ICY), were applied in core and outcrop examples to improve characterization of autocomposite ichnofabrics. Analyze Particles gives information about particle shape and size in the studied image. This tool was applied to Chondrites assemblages in composite ichnofabrics in view

Trace densities may be a potential proxy for tracemaker densities, especially in settings where traces but not body fossils preserve well. The relationship between the density of the gastropod Batillaria minima and its trails was examined in a modern muddy intertidal zone of a lagoon on San Salvador Island, the Bahamas. The number of snails found within a quadrat was a moderately positive predictor

The ichnospecies Gastrochaneolites dijugus Kelly and Bromley 1984 Kelly, S. R. A., and R. G. Bromley. 1984. “Ichnological Nomenclature of Clavate Borings.” Palaeontology 27: 793–807.[Web of Science ®] , [Google Scholar] and Teredolites longissimus Kelly and Bromley 1984 Kelly, S. R. A., and R. G. Bromley. 1984. “Ichnological Nomenclature of Clavate Borings.” Palaeontology 27: 793–807.[Web of Science

The ichnofossil Sanctum laurentiensis from the Late Ordovician (Cincinnatian) of Laurentia is a domichnium tunneled into live colonies of ramose and frondose trepostome Bryozoa. A preferred entrance location was chosen by the trace-making endoskeletozoan. The organism chose to tunnel in positions near growing tips of branches where one of two conditions prevailed. At stereotypical sites, here considered

The lower Maastrichtian deposits of the Mateur-Beja area in northern Tunisia are mainly composed of fine-grained marl and limestone alternations occasionally interbedded by coarse-grained calcarenites and gravel deposits. In the coarse-grained intervals sedimentary structures are indicative of storm-induced high-energy currents in an outer ramp to slope setting and of local reworking by bottom currents

Fluvial deposits of the uppermost Stockton Formation (Late Triassic), Newark Basin, west-central New Jersey have yielded an assemblage of trace fossils. Dominated by burrows, specimens include Cochlichnus anguineus, Helminthoidichnites tenuis, Planolites beverleyensis, Scoyenia gracilis, Spongeliomorpha carlsbergi, Treptichnus bifurcus, Treptichnus pollardi, plant remains, and an undetermined vertebrate

(2020). Book review. Ichnos: Vol. 27, No. 2, pp. 247-248.

The shallow continental shelf in the Cape Fear Region of southwestern Onslow Bay, North Carolina, contains lag deposits with an abundance of megatoothed shark teeth belonging to Otodus megalodon (Agassiz 1835) and Otodus chubutensis (Ameghino 1906) that derive from the Pliocene Yorktown and Miocene Pungo River formations, respectively. These teeth exhibit different frequencies and orientations of macroborings

Three parallel, manus-only sauropod trackways from the Coffee Hollow A-Male tracksite (Glen Rose Formation, Kendall County, Texas) were studied separately by researchers from the Heritage Museum of the Texas Hill Country and the Houston Museum of Natural Sciences. Footprint and trackway measurements generally show good agreement between the two groups’ data sets. Footprints appear to be shallowly impressed

This article presents a detailed description of insect trace fossils found in the megaflora of the Gzhelian-Asselian of the Bajo de Veliz. Plant-insect interactions are analyzed on 212 specimens of leaves from this flora, characterized by glossopterid (Gangamopteris, Euryphyllum, Glossopteris), cordaitales, sphenophytes, pteridosperms, lycophytes, conifers and seeds. Marginal feeding, surface and oviposition

The validity of the monotypic ichnogenus Thatchtelithichnus has been called into question due to an inference that it is a bioclaustration. Close analysis of the Thatchtelithichnus holmani holotype reveals that no part of this ichnotaxon involves a bioclaustration, and thus it remains a valid ichnotaxon. Thatchtelithichnus holmani has been observed on the bottom shell (plastron) of cryptodiran and

Sinusichnus Gibert 1996 Gibert, J. M. d. 1996. “A New Decapod Burrow System from the NW Mediterranean Pliocene.” Revista Española de Paleontología 11: 251–254. [Google Scholar] is recorded for the first time in the Oligocene-Miocene Pirabas Formation of Northern Brazil. In these Oligocene-Miocene carbonate deposits, Sinusichnus sinuosus is characterized by horizontal, highly regular sinusoidal burrow

When naming ichnotaxa based on uncollectable trace fossils, the holotype is the actual ichnofossil in the outcrop, though some ichnologists identify the holotype as a replica (cast) held in a museum collection, and refer to it as a “plastotype,” although not all such replicas are made from plaster. Nevertheless, through its Code, the International Commission on Zoological Nomenclature (ICZN) makes

(2020). Book review. Ichnos: Vol. 27, No. 2, pp. 244-246.

Ichnological analysis at the Pakhna Formation (Miocene, Cyprus) reveals, for the first time, the presence of graphoglyptid structures. The Pakhna Formation is dominated by pelagic/hemipelagic sediments, together with contourite, reworked turbidite and turbidite facies. Thus, a complex interaction between pelagic, bottom-current and gravitational sedimentation is envisaged. The discernment of facies

Marie Rouault was a French-born self-made paleontologist who left school at the age of 10. He then worked as a shepherd and spent a considerable portion of his life as a barber. During this period, he developed a keen interest in the rocks surrounding the city of Rennes in Brittany, northwestern France. All his free time was spent collecting the prolific fossils that he encountered. His collection

The ichnospecies Nummipera eocenica occurs in thick-bedded, tabular bioclastic massive sandstone of the upper Eocene San Jacinto Formation in the San Jacinto Fold Belt Basin, Colombia. Nummipera eocenica is a vertical to oblique burrow locally displaying a conical shape. The lining structure of the specimens studied consists of uniformly distributed lepidocyclinids and operculinids, with tests typically

In 1822, William Buckland first recognized Pleistocene vertebrate coprolites, and they are now known from more multiple localities with a global distribution. Carnivore coprolites dominate, and there are two distinct biogeographic and taphonomic provinces for vertebrate coprolites in the Pleistocene. The Castrocopros province of North and South America is characterized by a dominance of herbivore coprolites

A newly discovered track in the Chinle Group north of Moab, Utah, is attributable to the ichnogenus Atreipus, an ichnotaxon that is relatively common in eastern North America (Newark Supergroup) but very rare in the Late Triassic of the western part of the continent. This is only the second report of the genus from the Chinle Group. Atreipus has been attributed to a silesaurid dinosauriform, and dinosauriform

(2019). Obituary: Ron K. Pickerill (1947–2018) Ichnos: Vol. 26, No. 4, pp. 303-315.

This report documents the discovery of repichnia trace fossils Ptychoplasma (P. excelsum and P. vagans) and Dendroidichnites (D. irregulare); the fodichnia traces ?Ctenopholeus (?C. kutcheri) and cubichnia traces Bergaueria (B. hemishperica) from silty limestones of the Cretaceous Bagh Formation. These trace fossils have significant implications for the depositional facies and the paleo-environmental

The ichnogenus Kouphichnium and associated ichnofossils attributed to xiphosuran activity are here re-examined from samples collected from the Pennsylvanian-age Steven C. Minkin Fossil Site at the Union Chapel Mine, in Walker County, Alabama, USA. The large sample size offers an unique opportunity to evaluate some Kouphichnium ichnospecies. Thus, the morphological variability resulting from the taphonomic

Bioclast lined tubes of Schaubcylindrichnus coronus are not much reported around the world. As of now only two reports are available. Interestingly in both these reports and in the current find, tests of foraminifers are used as tube building material. Current report of Schaubcylindrichnus is from the subtidal deposits of the Naredi Formation (Ypresian) of Kachchh Basin, Western India. As the host

The tridactyl ichnotaxon Eubrontes giganteus commonly has been attributed to a carnivorous theropod dinosaur similar to Dilophosaurus or Liliensternus. For this to be correct, however, at least five unusual circumstances all must be true. (1) If the Eubrontes track-maker was a theropod, it created the most abundant large tracks found in the Connecticut Valley Hartford and Deerfield basins and yet,

Undichna quadrisulcata isp. nov. is described from the Eocene of Spitsbergen, where it occurs together with xiphosuran (horseshoe crabs) traces in upper shoreface to foreshore deltaic sandstone. This is the first record of a marine (deltaic) Undichna from the Eocene. The trail consists of two or three pairs of paralleling sinusoidal grooves (in epirelief) or ridges (in hyporelief). Previous records

The lower Paleozoic marine siliciclastic succession of the Central Andean Basin, northwestern Argentina, provides a valuable record of the onset of the Great Ordovician Biodiversification Event in western Gondwana. A new ichnospecies of rosette trace fossil, Gyrophyllites cristinae, is documented from lower and upper Tremadocian (Tr1 and Tr2) deposits of this basin. It is characterized by five to six

A new ichnospecies of Protopaleodictyon Książkiewicz, 1958 Książkiewicz, M. 1958. Stratigraphy of the Magura Series in the Średni Beskid (carpathians.). Biuletyn Instytutu Geologicznego 153:43–96. [In Polish, with English summary] [Google Scholar], Pr. aitkeni isp. n., is named from material recovered from the mid–Cambrian Stephen–Eldon formation transition in Banff National Park, Alberta, Canada.

The Yacoraite Formation (Maastrichtian–Danian; Balbuena Subgroup; Salta Group), from Northwestern Argentina, represents a shallow epeiric unit which is the result of transgressions in the Andean basin of South America. Herein, we study the avian footprints from the Maimará locality, Jujuy province, and Quebrada del Tapón ichnosite, Salta province. The avian footprints from the Maimará locality is less

The ichnogenus Radulichnus Voigt, 1977 Voigt, E. 1977. On grazing traces produced by the radula of fossil and recent gastropods and chitons. In Trace fossils 2, eds. T. P. Crimes, and J. C. Harper, 335–346. Geological Journal, Special Issue 9. [Google Scholar] is recorded for the first time from a bivalve, Anomalocardia brasiliana (Gmelin, 1791 Gmelin, J. F. 1791. Caroli a Linné, Systema Naturae. Tom

The fossil burrow Dolopichnus, capped by an ash layer, was collected ∼0.6 m below the Permian–Triassic Boundary at the Lung Cam section in North Vietnam. The burrow occurs at the top of Bed 9, directly below the stepped extinction in foraminifera that marks initiation of the global perturbations related to the end-Permian extinction event. Dolopichnus is a plug-shaped burrow, possessing lining and

Type specimens of the sauropod ichnotaxon Chuxiongpus changlingensis, which was later reassigned to Brontopodus changlingensis, as well as the theropod ichnotaxon Yunnanpus huangcaoensis, both from the Cretaceous Jiangdihe Formation of Yunnan Province, are redescribed in order to document their morphological features. Both, but particularly Y. huangcaoensis, which is considered now a nomen dubium,

A new ichnospecies of Cardioichnus, Cardioichnus biloba isp. nov., is described and documented from shallow marine strata of Cretaceous (Late Albian) age in southern New Mexico, USA. It is a heart-shaped resting trace which, unlike all other ichnospecies of Cardioichnus, is epichnial and increases in width to a bilobate trace, often with a raised anterior terminus. Only the posterior end of this trace

The aim of the present contribution is to describe large felipedid footprints from a new ichnological site from the Late Pleistocene of Buenos Aires Province, Argentina. The prints are referred as the new ichnospecies Felipeda miramarensis nov. ichnosp. Based on size and morphology, this new ichnotaxon may have been produced by the large machairodontine felid Smilodon populator. Track analysis indicates

(2018). First International Congress on Continental Ichnology (ICCI), El Jadida, Morocco, April 21–25, 2015. Ichnos: Vol. 25, First International Congress on Continental Ichnology (ICCI), El Jadida, Morocco, April 21-25, 2015, pp. 77-80.

Ichniotherium Pohlig, 1892 is a principal morphotype of Carboniferous-Permian tetrapod footprints referred to diadectomorphs. Though these tracks are relatively abundant in paleoequatorial regions of Pangea, the Cisuralian Bromacker locality in central Germany, is the only known place with co-occurring Ichniotherium ichnospecies. Ichniotherium cottae Pohlig, 1885 and Ichniotherium sphaerodactylum Pabst

(2018). Peter Pervesler (1.09.1951–25.10.2015) as an Ichnologist and Colleague. Ichnos: Vol. 25, No. 4, pp. 295-298.