The presence of trace fossils and their diversity in Miocene deposits of western Makran accretionary prism (SE Iran) was the center of this research. The trace fossil contents of Cheraghsuz and Sardasht (with Middle Miocene in age) and Gowharan (with Early Miocene in age) stratigraphic sections were analyzed. The recognized specimens of the Cheraghsuz section are Gordia isp., Helminthopsis hieroglyphica, Helminthopsis abeli, Helminthorhaphe flexsuosa, Paleodictyon maximum, and Spirorhaphe involuta. The Sardasht section is composed of Gordia isp., Helminthopsis hieroglyphica, Helminthopsis abeli, Helminthorhaphe flexsuosa, Paleodictyon maximum, Paleodictyon majus, Paleodictyon isp., Spirorhaphe involuta, Helicolithus isp., Helicorhaphe tortilis, and Neonereites isp.. In the Gowharan section, some gastropod trails, a giant and very well-preserved Archaeonassa-like ichnogenus, and ripple marks were observed. Majority of the recognized trace fossils are of the graphoglyptid group. The trace fossil contents of Cheraghsuz and Sardasht sections are compatible with Nereites ichnofacies; as graphoglyptids are thought to be indicative of oligotrophic conditions, the trace-producers are filter-feeder organisms that normally tend to take suspended nutrient materials in the water column, but due to lower sedimentation rate, the nutrient content is concentrated on the superficial part of the substrate; accordingly, the trace-makers prefer grazing and moving on the surface sediment to take their food. However, the exceptionally preserved trace fossils in Gowharan section should be formed in a tidal flat environment. In relation to the floor, it has recorded at least three shapes, including bi-lobed, uni-lobed and submerged forms. In addition, it could be observed that different trace fossils may be created by a single animal and different animals may create the same trace fossils.

中新世西部 Makran 增生棱柱（SE 伊朗）沉积物中的微量化石及其多样性是这项研究的中心。对 Cheraghsuz 和 Sardasht（年龄为中中新世）和 Gowharan（年龄为早中新世）地层剖面的微量化石含量进行了分析。Cheraghsuz 部分的公认标本是 Gordia isp .、Helminthopsis hieroglyphica、Helminthopsis abeli、Helminthorhaphe flexsuosa、Paleodictyon maximum和Spirorhaphe involuta。Sardasht 部分由Gordia isp.、Helminthopsis hieroglyphica、Helminthopsis abeli 组成, Helminthorhaphe flexsuosa , Paleodictyon maximum , Paleodictyon majus , Paleodictyon isp., Spirorhaphe involuta , Helicolithus isp., Helicorhaphe tortilis , and Neonereites isp..并观察到波纹痕迹。大多数公认的痕迹化石属于graphoglyptid组。Cheraghsuz 和 Sardasht 剖面的微量化石含量与 Nereites相容岩相; 由于石墨烯被认为是贫营养条件的指标，微量生产者是滤食性生物，通常倾向于在水柱中吸收悬浮的营养物质，但由于沉降率较低，营养成分集中在表层部分基材；因此，追踪者更喜欢在地表沉积物上放牧和移动来获取食物。但高华兰段保存异常完好的痕迹化石，应该是在滩涂环境中形成的。关于地板，它记录了至少三种形状，包括双叶形、单叶形和水下形状。此外，可以观察到，不同的痕迹化石可能由单一动物创造，不同的动物可能创造相同的痕迹化石。