Proterochampsidae are a group of predatory, putatively semi-aquatic, archosauriforms endemic to the Middle and Upper Triassic strata of Argentina and Brazil (Ischigualasto Formation and Santa Maria Supersequence). Here, the Brazilian species Proterochampsa nodosa from the Candelária Sequence (Santa Maria Supersequence of the Paraná Basin) is revisited with a detailed description, phylogenetic analysis and taxonomic revaluation. Through the application of computed tomographic scans, we were able to reveal part of the previously hidden morphology of its mandible and palate. We also discuss further differences between P. nodosa and P. barrionuevoi. The new phylogenetic analysis bolsters the monophyly of Proterochampsidae, the genus Proterochampsa and its position as the basalmost proterochampsid taxon in the clade.

中文翻译：

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坎德拉里亚序列（Santa Maria Supersequence）晚三叠世早期的 Proterochampsa nodosa（Archosauriformes：Proterochampsidae）的重新描述、分类重估和系统发育亲缘关系

Proterochampsidae 是阿根廷和巴西中上三叠统地层（伊斯基瓜拉斯托组和圣玛丽亚超层）特有的一组掠食性、推定为半水生的主龙形。在这里，对来自坎德拉里亚序列（巴拉那盆地的圣玛丽亚超序列）的巴西物种 Proterochampsa nodosa 进行了详细描述、系统发育分析和分类重估。通过计算机断层扫描的应用，我们能够揭示其下颌和上颚先前隐藏的部分形态。我们还讨论了 P. nodosa 和 P. barrionuevoi 之间的进一步差异。新的系统发育分析支持了 Proterochampsidae、Proterochampsa 属的单系及其作为进化枝中最基础的 proterochampsid 分类单元的地位。