Abstract Fish communities on tropical deep reefs are dominated by species that belong to families primarily composed of shallow-water species. Collections of deep-reef fishes via submersibles have allowed us to include these deep-reef species in molecular phylogenies, providing insights into the timing and frequency of invasions from shallow to deep reefs. Here we provide evidence of a new deep-reef invasion in the tribe Gobiosomatini in the family Gobiidae (gobies). We describe two new species, one of which belongs to a new genus, and incorporate these taxa into a time-calibrated molecular phylogeny of Gobiosomatini to show that, collectively, these two genera represent a previously unreported independent invasion on to deep reefs that occurred approximately 20–30 million years ago. These new taxa are readily distinguished from related genera and species by a combination of live coloration, pelvic-fin morphology, meristic characters, head-pore patterns and other osteological characters. We discuss the relevance of these two new species to the systematics of the tribe Gobiosomatini and include a comparison to all known genera in the tribe.

中文翻译：

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来自加勒比海的深礁虾虎鱼新谱系，包括两个新物种和一个新属（Teleostei: Gobiidae: Gobiosomatini）

摘要 热带深礁上的鱼类群落以属于主要由浅水物种组成的科的物种为主。通过潜水器收集深礁鱼类使我们能够将这些深礁物种纳入分子系统发育，从而深入了解从浅礁到深礁入侵的时间和频率。在这里，我们提供了 Gobiidae (gobies) 部落 Gobiosomatini 新的深礁入侵的证据。我们描述了两个新物种，其中一个属于一个新属，并将这些分类群纳入时间校准的 Gobiosomatini 分子系统发育，以表明，这两个属共同代表了以前未报道的对深礁的独立入侵，大约发生20-3000 万年前。这些新的分类群很容易通过活体颜色、腹鳍形态、分生特征、头孔模式和其他骨学特征的组合与相关属和种区分开来。我们讨论了这两个新物种与 Gobiosomatini 部落系统学的相关性，并包括与部落中所有已知属的比较。