We propose a simple biaxial tensional model which is able to reproduce at a qualitative level several aspects of early stages of vertebrate morphogenesis. The model is based on subsequent excitable contractions of an orthoradial and periclinal (radial) set of contracting lines, which generate first the basic embryonic pattern (a motile tube), and second the lateral orifices such as ears, eyes, mouth, gills, etc. An important aspect of the model is the self-arresting character of the process, akin to wound healing. At later stages, the biaxial lines may also work in extension, and this generates a developmental feedback which is quadratic with respect to curvature.

中文翻译：

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早期脊椎动物形态发生的双轴张力模型。

我们提出了一个简单的双轴张力模型，它能够在定性水平上再现脊椎动物形态发生早期阶段的几个方面。该模型基于一组正交和周（径向）收缩线的后续可兴奋收缩，这些收缩线首先生成基本胚胎模式（活动管），然后生成侧孔，例如耳朵、眼睛、嘴巴、鳃等. 该模型的一个重要方面是过程的自我抑制特性，类似于伤口愈合。在后期阶段，双轴线也可能在延伸中起作用，这会产生与曲率成二次的发育反馈。