Phylogenetic relationships of Vestini with extensive sampling were inferred using 37 taxa scored for 66 morphological characters. Consistent with recent DNA analyses, Vesta is found to be sister to Photurinae rendering Vestini monotypic. Conversely, our parsimony and model-based analyses robustly support the new subfamily Cladodinae established for Neotropical species of the former Vestini. The cladodine lineage includes Cladodes, Dodacles, Dryptelytra, Ledocas, plus the newly proposed Andecladodes gen. nov. and Brasilocladodes gen. nov. While Ledocas is mostly paraphyletic, Cladodes is polyphyletic and, therefore, Cladodes s.s. is introduced. Further, most trees presented a distinct Nyctocrepis, which is revalidated and includes the subgenus Fenestratocladodes (transferred from Cladodes). Finally, a highly supported crown clade designated as the Dodacles lineage is regularly recovered, splitting into two sister-clades: (1) Dodacles, Brasilocladodes and (2) Dryptelytra, C. proteus, C. solieri and Andecladodes. Four new Andean species are described: Andecladodes cosangensis sp. nov., A. ovalis sp. nov., Dryptelytra pampahermosae sp. nov. and Ledocas pikillactanus sp. nov. To recover the monophyly of genera, the following new combinations are proposed: Brasilocladodes carinatuscomb. nov., Brasilocladodes delalandeicomb. nov., Brasilocladodes illigericomb. nov., Dodacles lateraliscomb. nov. and Dodacles nigercomb. nov. (transferred from Cladodes); Ledocas emissuscomb. nov., Ledocas remixtuscomb. nov. (from Dodacles); Cladodes proteus comb. nov. (from Ledocas) Nyctocrepis malleri comb. nov. and Nyctocrepi stellatacomb. nov. (from Cladodes); and Cladodes cincticolliscomb. nov. and Cladodes melanuruscomb. nov. (from Vesta).

中文翻译：

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新亚科 Cladodinae 的系统发育：来自新热带地区的新生萤火虫（鞘翅目：Lampyridae）

使用对 66 个形态特征进行评分的 37 个分类群推断 Vestini 与广泛采样的系统发育关系。与最近的 DNA 分析一致，Vesta 被发现是 Photurinae 的姐妹，呈现 Vestini 单型。相反，我们的简约和基于模型的分析有力地支持为前 Vestini 的新热带物种建立的新亚科 Cladodinae。cladodine 谱系包括 Cladodes、Dodacles、Dryptelytra、Ledocas，以及新提出的 Andecladodes gen。十一月 和 Brasilocladodes gen。十一月 虽然 Ledocas 主要是并系的，但 Cladodes 是多系的，因此引入了 Cladodes ss。此外，大多数树木呈现出独特的 Nyctocrepis，它被重新验证并包括 Fenestratocladodes 亚属（从 Cladodes 转移）。最后，定期恢复被指定为 Dodacles 谱系的高度支持的皇冠进化枝，分裂成两个姐妹进化枝：（1） Dodacles、Brasilocladodes 和（2）Dryptelytra、C. proteus、C. solieri 和 Andecladodes。描述了四种新的安第斯物种：Andecladodes cosangensis sp。11 月，A. ovalis sp。十一月，Dryptelytra pampahermosae sp。十一月 和 Ledocas pikillactanus sp。十一月 为了恢复属的单一性，提出了以下新组合： Brasililocladodes carinatuscomb。十一月，Brasilocladodes delalandeicomb。十一月，Brasilocladodes illigericomb。11 月，Dodacles 侧梳。十一月 和 Dodacles nigercomb。十一月 （从 Cladodes 转移）；Ledocas emisuscomb。十一月，Ledocas remixtuscomb。十一月 （来自 Dodacles）；枝条变形梳。十一月 （来自 Ledocas）Nyctocrepis malleri 梳子。十一月 和 Nyctocrepi stellatacomb。十一月 （来自克拉多斯）；和枝条cincticcolliscomb。十一月 和 Cladodes melanuruscomb。十一月 （来自灶神星）。