In fleshy fruits, organic acids are the main source of fruit acidity and play an important role in regulating osmotic pressure, pH homeostasis, stress resistance, and fruit quality. The transport of organic acids from the cytosol to the vacuole and their storage are complex processes. A large number of transporters carry organic acids from the cytosol to the vacuole with the assistance of various proton pumps and enzymes. However, much remains to be explored regarding the vacuolar transport mechanism of organic acids as well as the substances involved and their association. In this review, recent advances in the vacuolar transport mechanism of organic acids in plants are summarized from the perspectives of transporters, channels, proton pumps, and upstream regulators to better understand the complex regulatory networks involved in fruit acid formation.

中文翻译：

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植物液泡中有机酸积累的机制与调控

在肉质果实中，有机酸是果实酸度的主要来源，在调节渗透压、pH 稳态、抗逆性和果实品质等方面发挥着重要作用。有机酸从胞质溶胶到液泡的运输及其储存是复杂的过程。在各种质子泵和酶的帮助下，大量的转运蛋白将有机酸从胞质溶胶运送到液泡。然而，关于有机酸的液泡运输机制以及所涉及的物质及其关联，仍有许多待探索。本文从转运体、通道、质子泵、