The cosmetids are conspicuous harvestmen, remarkably diverse in size, shape and colour. However, the effectiveness of all these morphological traits for diagnosing groups is scarcely explored in the literature. Since the early 20th century, the family Cosmetidae has been divided into two subfamilies, Cosmetinae and Discosomaticinae, and there has been no further effort to delimit additional or alternative subfamilial groups. In this analysis, we aim to test the issue of the basal dichotomy of Cosmetinae and Discosomaticinae. Thus, we propose a more comprehensive phylogenetic analysis using both parsimony and Bayesian approach, comprising 103 terminals of Cosmetidae, plus seven outgroup terminals scored for 130 morphological characters. Discosomaticinae is revisited and all its species were included in our matrix. To offer a real challenge to the monophyly of Discosomaticinae, members of 36 genera of Cosmetinae have also been scored. Our results support neither Cosmetinae nor Discosomaticinae sensu Pickard-Cambridge/Roewer as monophyletic groups. We found that Cosmetidae may be organized into a few major clades, which are here diagnosed. Accordingly, taxonomic changes in the arrangement of the family are introduced: description of one new tribe, five new subfamilies and revalidation of a sixth, description of three new genera, and proposition of synonymies, transfers and revalidations at generic level.

中文翻译：

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基于形态学的分支学分裂了百年历史的花斑收割者二分法（蛛形纲：Gonyleptoidea：Cosmetidae）

cosmetids 是显眼的收获者，在大小、形状和颜色上非常多样化。然而，文献中几乎没有探讨所有这些形态特征对诊断群体的有效性。自 20 世纪初以来，Cosmetidae 家族已分为两个亚科，Cosmetinae 和 Discosomaticinae，并且没有进一步努力划定额外或替代的亚科群。在这项分析中，我们旨在测试 Cosmetinae 和 Discosomaticinae 的基础二分法问题。因此，我们提出使用简约法和贝叶斯方法进行更全面的系统发育分析，包括 103 个 Cosmetidae 末端，以及 7 个外群末端，对 130 个形态特征进行评分。Discosomaticinae 被重新审视，它的所有物种都包含在我们的矩阵中。为了对盘螈的单一系提出真正的挑战，还对 36 属的 Cosmetinae 成员进行了评分。我们的结果既不支持 Cosmetinae 也不支持 Discosomaticinae sensu Pickard-Cambridge/Roewer 作为单系群。我们发现 Cosmetidae 可能被组织成几个主要的进化枝，这些进化枝在这里被诊断出来。相应地，介绍了科排列的分类学变化：一个新部落的描述，五个新的亚科和第​​六个的重新验证，三个新属的描述，以及同义词的命题，转移和重新验证。我们发现 Cosmetidae 可能被组织成几个主要的进化枝，这些进化枝在这里被诊断出来。相应地，介绍了科排列的分类学变化：一个新部落的描述，五个新的亚科和第​​六个的重新验证，三个新属的描述，以及同义词的命题，转移和重新验证。我们发现 Cosmetidae 可能被组织成几个主要的进化枝，这些进化枝在这里被诊断出来。相应地，介绍了科排列的分类学变化：一个新部落的描述，五个新的亚科和第​​六个的重新验证，三个新属的描述，以及同义词的命题，转移和重新验证。