Heartwood colour is often an important factor in determining timber prices. However, the determinants of intraspecific variation in heartwood colour, which is useful information for sustainable wood marketing, are little understood, especially at the local scale in cool temperate forests. Because heartwood is produced as a secondary compound and photosynthesis is regulated by nitrogen (N) in cool temperate forests, we hypothesized that (1) soil conditions determine heartwood colour even at a local scale within a tree species and (2) N, specifically, can be an important driver of the intraspecific variation in heartwood colour in the trees of cool temperate forests. To test these hypotheses, we investigated the relationship between the colour values (luminescence, redness, and yellowness) of heartwood from Juglans mandshurica var. sachalinensis and the soil parameters in a cool temperate forest. Among the soil properties, not soil N but soil magnesium (Mg) contents alone had a significant influence on the redness and yellowness of the heartwood. Higher soil Mg contents resulted in increased redness and yellowness of the heartwood in our study, probably due to the increase in phenolics and the colouring of the tannins in the heartwood with Mg. Our results indicate that even at a local scale, soil condition can determine the intraspecific variation in heartwood colour and that forest managers can utilize edaphic information to predict heartwood colour for timber marketing.

中文翻译：

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土壤性质对胡桃心材颜色的影响。sachalinensis在凉爽的温带森林中

心材颜色通常是决定木材价格的重要因素。然而，对于可持续木材营销有用的信息，心材颜色种内变化的决定因素知之甚少，尤其是在凉爽温带森林的局部范围内。因为心材是作为次生化合物生产的，并且光合作用在凉爽温带森林中受氮 (N) 调节，我们假设 (1) 土壤条件决定了心材的颜色，即使在树种的局部范围内，(2) N，具体来说，可能是冷温带森林树木心材颜色种内变化的重要驱动因素。为了验证这些假设，我们研究了胡桃心材的颜色值（发光、红色和黄色）之间的关系。sachalinensis 和凉爽温带森林中的土壤参数。在土壤特性中，不是土壤 N 而是土壤镁 (Mg) 含量单独对心材的红色和黄色有显着影响。在我们的研究中，较高的土壤镁含量导致心材发红和发黄的程度增加，这可能是由于酚类物质的增加和心材中单宁的着色与镁有关。我们的结果表明，即使在局部范围内，土壤条件也可以确定心材颜色的种内变化，并且森林管理者可以利用土壤信息来预测木材营销的心材颜色。在我们的研究中，较高的土壤镁含量导致心材发红和发黄的程度增加，这可能是由于酚类物质的增加和心材中单宁的着色与镁有关。我们的结果表明，即使在局部范围内，土壤条件也可以确定心材颜色的种内变化，并且森林管理者可以利用土壤信息来预测木材营销的心材颜色。在我们的研究中，较高的土壤镁含量导致心材发红和发黄的程度增加，这可能是由于酚类物质的增加和心材中单宁的着色与镁有关。我们的结果表明，即使在局部范围内，土壤条件也可以确定心材颜色的种内变化，并且森林管理者可以利用土壤信息来预测木材营销的心材颜色。