The precinctive New Caledonian genus Cyphocoleus Chaudoir is revised with 22 species recognized, 12 newly described: C. lissus sp. n., C. prolixus sp. n., C. parovicollis sp. n., C. burwelli sp. n., C. angustatus sp. n., C. monteithi sp. n., C. fasciatus sp. n., C. lescheni Liebherr & Will, sp. n., C. cordatus sp. n., C. bourailensis sp. n., C. subulatus sp. n., and C. iledespinsensis sp. n. Atongolium Park & Will is found to be a junior synonym of Cyphocoleus, with its two species recombined as C. mirabilis comb. n. and C. moorei comb. n. Results of a survey of Harpalinae Bonelli place Cyphocoleus as a member of Odacanthini based on synapomorphies of the eighth abdominal tergite and the female spermathecal assembly. Cyphocoleus shares with five other generic-level taxa – Homethes Newman, Aeolodermus Andrewes, Stenocheila Laporte, Quammenis Erwin and Diplacanthogaster Liebke – a single-segmented maxillary galea that is appressed to the outer margin of the maxillary lacinia. These six generic-level taxa are newly classified as members of subtribe Homethina subtrib. n. (type genus Homethes). Cladistic analysis including 79 taxa and utilizing 119 morphological characters supports division of Odacanthini into four monophyletic subtribes: 1, Actenonycina (Actenonyx White); 2, Homethina; 3, Pentagonicina (Pentagonica Dana, Parascopodes Darlington, Scopodes Erichson); and 4, Odacanthina (24 genera in this analysis monophyletically defined by Lasiocera Dejean and its adelphotaxon). These subtribes are phylogenetically arranged as: (Actenonycina (Homethina (Pentagonicina + Odacanthina). Area relationships defined within Homethina – (New Caledonia (Australia (South America + Central America))) – support the origin of New Caledonian Cyphocoleus prior to amphiantarctic vicariance between South America and Australia. Consistent with previous molecular dating of 100–105 Ma for the origin of Odacanthini, a general vicariance-based hypothesis proposes that New Zealandian Actenonyx and New Caledonian Cyphocoleus were emplaced on Zealandia prior to the completion of rifting between Zealandia and Australia during Late Cretaceous, and that both fragments of Zealandia remained subaerial throughout the Cenozoic. Alternatively, under a very specific time-constrained biogeographic hypothesis ladened with an added assumption of dispersal, the ancestor of Cyphocoleus could have colonized New Caledonia during a 2–5 Ma period after its proposed subaerial reemergence at 37 Ma. A clade within Cyphocoleus synapomorphously exhibits an environmental patina: a varnish-like coating to the dorsal body surface that is hypothesized to enable crypsis of the adult beetle. Several specializations of elytral setae are also synapomorphies of this clade, suggesting evolutionary association of the patina and the setal specializations. Key Words

中文翻译：

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Cyphocoleus Chaudoir（鞘翅目、Carabidae、Odacanthini）：描述性分类、系统发育关系和新喀里多尼亚的新生代历史

辖区的新喀里多尼亚属 Cyphocoleus Chaudoir 进行了修订，确认了 22 个物种，新描述了 12 个物种：C. lissus sp.。n., C. prolixus sp. n., C. parovicolis sp. n., C. burwelli sp. n., C. angustatus sp. n., C. monteithi sp. n., C. fasciatus sp. n., C. lescheni Liebherr & Will, sp. n., C.cordatus sp. n., C. borailensis sp. n., C. subulatus sp. n. 和 C. iledespinsensis sp. n. Atongolium Park & Will 被发现是 Cyphocoleus 的次要同义词，它的两个物种重组为 C. mirabilis 梳子。n. 和 C. moorei 梳子。n. 对 Harpalinae Bonelli 的调查结果根据第八腹部 tergite 和雌性受精囊组件的突触形态将 Cyphocoleus 列为 Odacanthini 的成员。Cyphocoleus 与其他五个通用级分类群共享——Homethes Newman、Aeolodermus Andrewes、Stenocheila Laporte、Quammenis Erwin 和 Diplacanthogaster Liebke – 一种单节的上颌盖帽，贴附在上颌裂孔的外缘。这六个属级分类群被新分类为亚部落 Homethina 亚部落的成员。n. （类型属 Homethes）。包括79个分类群和利用119个形态特征的分支分析支持将Odacanthini分为四个单系亚部落：1，Actenonycina（Actenonyx White）；2、家居；3、五角星（Pentagonica Dana、Parascopodes Darlington、Scopodes Erichson）；和 4，Odacanthina（本分析中的 24 个属由 Lasiocera Dejean 及其 adelphotaxon 单系定义）。这些亚部落在系统发育上排列为：（Actenonycina (Homethina (Pentagonicina + Odacanthina)）。Homethina 内定义的区域关系（新喀里多尼亚（澳大利亚（南美洲 + 中美洲）））支持新喀里多尼亚 Cyphocoleus 在南美洲和澳大利亚之间的两极替代之前的起源。与之前关于 Odacanthini 起源的 100-105 Ma 分子测年一致，一个基于一般替代性的假设提出，新西兰 Actenonyx 和新喀里多尼亚 Cyphocoleus 在晚白垩世西兰洲和澳大利亚之间的裂谷完成之前就已经安置在西兰洲上，并且在整个新生代中，西兰西亚的两个碎片都保持在地下。或者，在一个非常具体的时间受限的生物地理假设下，加上一个额外的扩散假设，Cyphocoleus 的祖先可能在其提议的 37 Ma 地下重新出现后的 2-5 Ma 期间在新喀里多尼亚殖民。Cyphocoleus 内的一个进化枝突触地表现出环境铜绿：背部体表的一层清漆状涂层，被假设为使成年甲虫能够隐匿。鞘翅刚毛的几个特化也是这个进化枝的突触，表明古色和刚毛特化的进化关联。关键词 建议 patina 和 setal 专业化的进化关联。关键词 建议 patina 和 setal 专业化的进化关联。关键词