The functional response of natural patches to surrounding land-use changes is strongly related to variations in functional traits of coexisting species. To exemplify the effects on species of a general pattern of land-use intensification mountains-coastland, we investigated the variation of a key plant trait - seed mass - in small woodlots located along a land-use intensification gradient for two common species (Asparagus albus and Asparagus acutifolius) in the Mediterranean areas. Moreover, along this gradient, we also explored the relationship between seed mass variation and different environmental filters.

Along the gradient, A. acutifolius seed mass decreased from natural and semi-natural to urban and artificial areas (higher to lower elevation), whereas A. albus seed mass increased along the same gradient, with heavier seed in patches located in the urban and artificial areas than in those located in natural and semi-natural areas. At intra-specific level, A. acutifolius seeds were significantly different at the extremes of the gradient (natural and semi-natural vs urban and artificial areas), while A. albus showed significant differences both between natural and semi-natural areas and urban and artificial areas, and between agricultural and urban and artificial areas, revealing more sensitiveness to land-use change.

The land-use type influenced seed mass variability: in the small patches located in natural and semi-natural areas and in agricultural ones, we observed for both species a higher seed mass variability, being highest in the agricultural areas, while we observed a limited variability in urban and artificial areas, suggesting a homogenization in terms of seed mass within and across species in human-altered areas. Environmental drivers on the seed mass of the two species showed an opposite trend in relation to biotic, topographic and bioclimatic variables.

We observed that for two common Mediterranean species, land-use type influenced one of the most important plant functional traits (i.e., seed mass), leading to a reduction of intraspecific variability in artificial context. Understanding how and why these relations occur could improve our capacity to find adaptive strategies for environmental management.

自然斑块对周围土地利用变化的功能响应与共存物种功能性状的变化密切相关。为了举例说明土地利用集约化山脉 - 海岸的一般模式对物种的影响，我们研究了两个常见物种（白芦笋）在位于土地利用集约化梯度上的小林地中的关键植物性状 - 种子质量 - 的变化和芦笋 acutifolius）在地中海地区。此外，沿着这个梯度，我们还探索了种子质量变化与不同环境过滤器之间的关系。

沿着梯度，A. acutifolius种子质量从自然和半自然到城市和人工区域（海拔从高到低）减少，而A. albus种子质量沿着相同的梯度增加，位于城市和人工区域比位于自然和半自然区域的区域。在种内水平上，A. acutifolius种子在梯度的极端（自然和半自然与城市和人工区域）显着不同，而A. albus在自然和半自然区域以及城市和人工区域之间显示出显着差异。人工区域，以及农业与城市与人工区域之间，显示出对土地利用变化更加敏感。

土地利用类型影响种子质量变异性：在位于自然和半自然区域以及农业区域的小块中，我们观察到两个物种的种子质量变异性更高，在农业区最高，而我们观察到的有限城市和人工区域的变异性，表明在人类改变的地区，物种内部和物种之间的种子质量存在同质化。两个物种种子质量的环境驱动因素在生物、地形和生物气候变量方面表现出相反的趋势。

我们观察到，对于两种常见的地中海物种，土地利用类型影响了最重要的植物功能性状之一（即种子质量），从而减少了人工环境中的种内变异性。了解这些关系如何以及为何发生可以提高我们寻找环境管理适应性策略的能力。