Non-structural carbohydrates (NSC) reserves are crucial for trees to cope with weather extremes, thus to ensure their survival and ecological plasticity. The NSC reserves can depend on social status, suggesting uneven plasticity of trees at the stand level. In stemwood of Scots pine (Pinus sylvestris L.), which is a widespread and important species, NSC reserves are stored in parenchyma in wood rays (WR). The quantity of WRs is adjusted intra-annually, allowing retrospective analysis of factors affecting their formation. Accordingly, the differences in WR quantity in stemwood of dominant and intermediate (canopy trees with reduced and narrow crowns) maturing Scots pine were assessed by quantitative wood anatomy. Tangential cuts from the outermost 30 tree-rings were analysed. The relative ray area was intermediate, i.e. covering about 5% of the tangential cut, yet expressed high individuality among the trees. The size and amount of WR mainly differed between the earlywood and latewood; WRs in latewood were higher although narrower in comparison to earlywood, yet their total amount was higher in earlywood. Canopy status had only a slight effect, as quantity and height of WR tended to be higher for the intermediate trees, particularly in earlywood. The size and quantity of WR expressed inter-annual variation, which was mainly related to the meteorological conditions prior to the formation of the tree-ring (previous summer and autumn) indicating legacy effects of climatic factors on NSC and susceptibility of trees to cumulative effects of weather extremes. However, the climatic signals in the inter-annual variation of WR were weaker than observed before, likely due to location of the studied stand in the mid-part of the species range. Nevertheless, the observed differences in mean values and inter-annual variation of WR suggested a within-species diversity of carbon allocation patterns, supporting adaptability of the species.

非结构性碳水化合物（NSC）储备对于树木应对极端天气至关重要，从而确保树木的生存和生态可塑性。NSC的储备可能取决于社会地位，这表明林分水平上树木的可塑性不均匀。在苏格兰松木（樟子松L.）是一种广泛而重要的物种，NSC储藏在木射线（WR）的薄壁组织中。WR的数量每年调整一次，从而可以回顾性分析影响其形成的因素。因此，通过定量木材解剖学评估了成熟的苏格兰松树的主要和中间（冠冠减少且冠状狭窄的冠层树）的树干木材中WR量的差异。分析了从最外面的30个年轮的切向切割。相对射线区域是中间的，即覆盖约5％的切向切口，但在树木之间表现出很高的个性。早熟木材和晚熟木材的WR大小和数量主要不同。与早材相比，晚材中的WR较高，但较窄，但早材中的WR总量较高。机盖状态仅产生轻微影响，因为中间树木的WR数量和高度往往更高，尤其是早木。WR的大小和数量表示年际变化，这主要与树木年轮形成之前的气象条件（夏季和秋季）有关，这表明气候因素对NSC的遗留影响以及树木对累积影响的敏感性极端天气。但是，WR的年际变化中的气候信号比以前观测到的要弱，这可能是由于研究林分在物种范围的中部。然而，观察到的WR平均值和年际变化的差异表明物种内部的碳分配模式具有多样性，从而支持了物种的适应性。