With more than 300 extant species, Frullaniaceae represent a species-rich clade of the predominantly epiphytic order Porellales, occurring in humid tropical and subtropical forests as well as temperate regions. Earliest fossils of Frullaniaceae are known from mid-Cretaceous Burmese and Alaskan ambers. So far, evidence for four or five fossil species have been recovered in these ambers. The steady search for new fossils is important, since many liverwort fossils lack important characters and sometimes only the investigation of numerous specimens can provide all relevant characters. Based on two newly discovered and exquisitely preserved gametophytes from mid-Cretaceous Burmese amber, we provide a detailed investigation and re-appraisal of two previously described species of Frullaniaceae, namely Frullania baerlocheri and Protofrullania cornigera. The new discovery of a complete gametophyte of F. baerlocheri with both vegetative and fertile parts enables an emended morphological description, and demonstrates that F. baerlocheri and F. pinnata are conspecific. A new specimen of P. cornigera provides additional information concerning leaf lobule, stylus and leaf cells. The fossil record and phylogenetic analysis of the liverworts recovered in mid-Cretaceous Burmese and Alaskan amber clearly show that Frullaniaceae have already accumulated some notable diversity in the Cretaceous, including the extant genus Frullania with three species and the extinct monotypic genus Protofrullania from mid-Cretaceous Burmese amber and probably also the extinct monotypic genus Kaolakia from mid-Cretaceous Alaskan amber.

Frullaniaceae拥有300多种现存物种，代表着一个主要为附生类Porellales的物种丰富的进化枝，分布于潮湿的热带和亚热带森林以及温带地区。Frullaniaceae的化石最早是从白垩纪中期的缅甸和阿拉斯加的琥珀中得知的。到目前为止，已经在这些琥珀中找到了四到五个化石物种的证据。不断寻找新的化石很重要，因为许多地蒿化石缺乏重要的特征，有时只有对大量标本进行调查才能提供所有相关的特征。基于白垩纪中期缅甸琥珀的两种新近发现且保存完好的配子体，我们对先前描述的两种菊苣菜科植物菊苣菜进行了详细的调查和重新评估。和Protofrullania cornigera。F完整配子体的新发现。baerlocheri与两个无性和肥沃部件使得能够仔细的校勘形态的说明中，并表明˚F。baerlocheri和˚F。Pinnata是同种的。P的一个新样本。Cornigera提供了有关叶小叶，触笔和叶细胞的其他信息。化石记录和对在白垩纪中期缅甸和阿拉斯加琥珀中回收的艾蒿的系统发育分析清楚地表明，菊苣科已经在白垩纪积累了一些显着的多样性，包括现存的属。耳叶有三个品种和已灭绝的单种属Protofrullania从白垩纪中期缅甸琥珀，也可能已灭绝的单种属Kaolakia从白垩纪中期阿拉斯加琥珀色。