Direct multiyear observations were carried out to track changes in the species composition and structure of extrazonal steppes in the boreal zone of the Southern Urals inhabited by many rare plant species. It is shown that the mesophytization of communities has taken place in petrophytic steppes in the period of 1999–2019: the cover of meadow–steppe and steppe species increased by 6.8 times; while the cover of petrophytic-steppe species, by only 2.2 times. A portion of petrophytic steppes has been absorbed by surrounding meadow steppes. It is established that communities less than 40 m2 in size are most susceptible to the changes. The observed changes are likely caused by an increase in precipitation in the beginning of the growing season.

中文翻译：

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气候变化下 20 年来南乌拉尔带外草原的动态

进行了多年的直接观测，以追踪栖息着许多稀有植物物种的南乌拉尔北部地区带外草原的物种组成和结构的变化。结果表明，1999-2019年岩性草原群落发生了中生化：草甸草原和草原物种覆盖率增加了6.8倍；而岩生草原物种的覆盖率仅为2.2倍。一部分岩质草原已被周围的草甸草原吸收。已确定小于 40 平方米的社区最容易受到这些变化的影响。观察到的变化很可能是由于生长季节开始时降水量的增加造成的。