Abstract During the Permian, major icehouse-greenhouse climate shifts and tectonic reconfiguration had important biogeographic implications, especially for climate-sensitive organisms such as fusulinids. Here we present multivariate methods on a global fusulinid species dataset including 1546 species from 58 localities in the Early (Asselian, Sakmarian, Artinskian and Kungurian) and Middle (Roadian, Wordian and Capitanian) Permian. Our results show that fusulinid global provincialism was high in the Asselian, Sakmarian, and Artinskian, driven by the development of multiple fusulinid bioregions in and near the Tethys Ocean. During the Asselian, Uralian sites and nearby regions of western Tethys were distinct from eastern Tethys, while stations in Arctic Russia and Norway formed a separate Boreal bioregion. Tectonic closure of the oceanic gateway in the southern Urals resulted in progressive isolation of the Uralian and Boreal bioregions during the Sakmarian and Artinskian and their ultimate disappearance by the Kungurian. Climate warming likely was the most important control on the Sakmarian formation of the distinct peri-Gondwana bioregion, because its development coincided with deglaciation following the Late Paleozoic Ice Age but preceded the separation of the Cimmerian terranes from northern margin of Gondwana. On the other hand, northward movement of the Cimmerian blocks following Artinskian-Kungurian rifting ultimately led to the merger of the peri-Gondwanan bioregion with tropical Tethyan faunas, resulting in lower provincialism in the Guadalupian and minimal faunal differentiation across Tethys. In contrast, faunal similarity between Tethys and eastern Panthalassa (the McCloud region and southwestern United States) was higher in the Asselian-Artinskian but decreased in the Kungurian and Middle Permian, perhaps as the result of sluggish ocean circulation following the warming episode of Late Paleozoic deglaciation.

中文翻译：

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Cisuralian 和 Guadalupian 全球古生物地理学对构造、海洋环流和气候变化的响应

摘要 在二叠纪期间，主要的冰库-温室气候变化和构造重构具有重要的生物地理意义，尤其是对气候敏感的生物，如梭菌。在这里，我们介绍了全球 Fusulinid 物种数据集的多变量方法，包括来自早 (Asselian、Sakmarian、Artinskian 和 Kungurian) 和中 (Roadian、Wordian 和 Capitanian) 二叠纪 58 个地点的 1546 种物种。我们的研究结果表明，由于特提斯洋内和附近多个梭生物区的发展，在 Asselian、Sakmarian 和 Artinskian 中，fusulinid 全球地方性很高。在阿瑟利亚时期，乌拉尔遗址和特提斯西部附近地区与特提斯东部不同，而北极俄罗斯和挪威的站点形成了一个单独的北方生物区。乌拉尔南部海洋门户的构造关闭导致乌拉尔和北方生物区在萨克马和阿廷斯基期逐渐隔离，最终被昆古里亚人消失。气候变暖可能是对独特的冈瓦纳周围生物区萨克马里亚形成的最重要的控制，因为它的发展与晚古生代冰河时代之后的冰川消融同时发生，但在辛梅里地体与冈瓦纳北缘分离之前。另一方面，在阿廷斯基-孔古尔裂谷之后，辛梅里亚地块向北移动，最终导致了冈瓦纳周围生物区与热带特提斯动物群的合并，导致瓜达卢皮亚的地方性降低，特提斯的动物区系分化最小。相比之下，