A new subgenus of Tenebroides, Polynesibroides subgen. nov., is established for six potentially extinct beetles, Tenebroides atiu, T. moorea, T. raivavae, T. rimatara, T. tubuai and T. mihiura spp. nov., described from the Cook, Austral and Society Islands based on subfossil fragments, and the extant T. tahiti sp. nov. from the island of Tahiti, Society Islands. Here we present the first detailed time-scaled phylogeny of the superfamily Cleroidea inferred from our four-gene dataset, including T. tahiti. Bayesian tip-dating, incorporating 17 fossils, estimated that Cleroidea appeared at the end of the Triassic (~210 Mya). The split of Biphyllidae + Byturidae + Phloiophilidae was reconstructed at ~204 Mya, followed by Early Jurassic separation of Trogossitidae (~200 Mya), Acanthocnemidae (~197 Mya), stem Cleridae–Thanerocleridae–Chaetosomatidae (~194 Mya), Peltidae, Lophocateridae, Decamerinae and the melyrid lineage (~191 Mya). Trogossitidae diversified by separation of Kolibacia (~136 Mya) in the Cretaceous, followed by parting Temnoscheila + Nemozoma and Airora + Tenebroides lineages (~113 Mya). Tenebroides radiated ~75 Mya and T. tahiti diverged from Panamanian taxa in the Middle Eocene (~40 Mya). Fourteen morphological characters of Pacific trogossitids were analysed. Parallelodera, including the Panamanian species, is nested within Tenebroides being sister to Polynesibroides. Therefore, Parallelodera is classified as a subgenus of Tenebroides. Tenebroides fairmairei is placed in the subgenus Parallelodera. A Neotropical origin for Parallelodera and Polynesibroides is inferred.

中文翻译：

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晚期三叠纪生物危机后灭绝和现存的太平洋蛇口科和鞘翅目（鞘翅目）的演变

Tenebroides的一个新亚种，Polynesibroides亚种。十一月 建立了用于六个可能灭绝的甲虫的Tenebroides atiu，T。moorea，T。raivavae，T。rimatara，T.tubuai和T. mihiura spp。11月的，来自库克，澳和基于亚化石片段社会群岛，和现存的描述T.塔希提岛藻。十一月 来自社会群岛的塔希提岛。在这里，我们展示了从我们的四基因数据集（包括塔希提岛）推断出的超家族克氏杆菌的第一个详细的时标系统发育史。。贝叶斯尖端约会，结合了17种化石，估计Cleroidea出现在三叠纪末期（〜210 Mya）。在〜204 Mya处重建了Biphyllidae + Byturidae + Phloiophilidae的分裂，然后对侏罗纪的Trogossitidae（〜200 Mya），Acanthocnemidae（〜197 Mya），Cleridae-Thanerocleridae-Chaetosomatidae（〜194 Mya），Peltidae， ，Decamerinae和子粒系（〜191 Mya）。Trogossitidae通过在白垩纪分离Kolibacia（〜136 Mya）多样化，然后分开Temnoscheila + Nemozoma和Airora + Tenebroides世系（〜113 Mya）。Tenebroides辐射〜75 Mya和T.tahiti在中始新世（〜40 Mya）中脱离了巴拿马的生物分类。分析了太平洋蛇的十四个形态特征。Parallelodera，包括巴拿马物种，被嵌套在Tenebroides内，是Polynesibroides的姐妹。因此，Parallelodera被归类为Tenebroides的亚属。Tenebroides fairmairei放在Parallelodera属下。对于A新热带起源Parallelodera和Polynesibroides推断。