Eocene vertebrate footprints from the uppermost layers of the Karaj Formation have been discovered in the Taleghan valley, Alborz Mountains, northern Iran. Trackways comprise carnivore and artiodactyl footprints. Footprints of primitive and small artiodactyl are assigned to Gambapes hastatus, with 16–18 mm footprint length. Carnivore trackways belong to the Felidae or Mesonychidae, recognized as Pycnodactylopus cf. achras. These footprints and previous reports of track sites from the Alborz Mountains document a high diversity of Eocene track-makers in this region of the Iranian Plateau.

中文翻译：

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伊朗北部阿尔伯兹山脉塔莱汉谷地区卡拉伊组（始新世）的食肉动物和偶蹄动物哺乳动物的足迹

在伊朗北部Alborz山的Taleghan谷地发现了Karaj组最上层的始新世脊椎动物足迹。轨道包括食肉动物和偶蹄动物足迹。原始的和小的偶蹄动物的脚印分配给了Gambapes hastatus，足迹长度为16-18 mm。食肉动物trackways属于猫科或中爪兽科，公认Pycnodactylopus比照 阿克拉斯。这些足迹和先前来自Alborz山脉的跟踪站点的报告表明，伊朗高原这一地区的始新世跟踪器种类繁多。