Species of Verrucaria, characterised by large spores (at least some spores exceeding 25 µm in length), perithecia leaving pits in the rock and a pale thin thallus, form a taxonomically-difficult and poorly-known group. In this study, such species occurring in Finland are revised, based on ITS sequences and morphology. Maximum likelihood analysis of ITS sequence data was used to examine if the species belong to the Thelidium group, as suggested by BLAST search. Twelve species are accepted in Finland: Verrucaria bifurcatasp. nov., V. cavernarumsp. nov., V. devergens, V. difficilissp. nov., V. foveolata, V. fuscozonatasp. nov., V. karelica, V. kuusamoensissp. nov., V. subdevergenssp. nov., V. subjunctiva, V. subtilis and V. vacillanssp. nov.Verrucaria foveolata is nested in V. subjunctiva in the phylogeny, but due to morphological and ecogeographical differences, the two taxa are treated as separate species pending further studies. Based on the analysis, the study species belong to the Thelidium group. The studied species show a rather high infraspecific morphological, but a low genetic variation. Furthermore, they show considerable overlap in their morphology and many specimens cannot be reliably identified, based on morphology only. All species are restricted to calcareous rocks. Verrucaria alpigena, V. cinereorufa and V. hochstetteri are excluded from the lichen flora of Finland. Verrucaria grossa is considered a species with unresolved identity. Verrucaria foveolata and V. subtilis are rather common on calcareous rocks of Finland while V. devergens and V. kuusamoensis are restricted to northern Finland. Verrucaria subjunctiva occurs mainly in northern Finland. Verrucaria bifurcata has been found only from southern Finland. Verrucaria difficilis has few localities both in SW and NE Finland. Verrucaria vacillans is restricted to calcareous rocks (dolomite) on the mountains of the NW corner of Finland. Verrucaria fuscozonata, V. karelica and V. subdevergens occur only in the Oulanka area in NE Finland. A lectotype is designated for V. subjunctiva. The morphology of the Finnish species was compared with 51 European species of Verrucaria presumably belonging to the Thelidium group.

中文翻译：

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Verrucaria种类的分类法，其特征是大孢子，蠕虫皮在岩石中留下凹坑，而芬兰则是浅薄的all。

Verrucaria的特征是大的孢子（至少一些孢子的长度超过25 µm），蠕虫皮在岩石中留下小凹坑和浅薄的all肉，这在分类学上是很难分类的，并且鲜为人知。在这项研究中，基于ITS序列和形态对在芬兰发生的此类物种进行了修订。如BLAST搜索所建议的，使用ITS序列数据的最大似然分析来检查该物种是否属于Thelidium组。芬兰接受了十二种：Verrucaria bifurcatasp。11月，V。Cavernarumsp。11月，V。devergens，V。difficilissp。11月，V。foveolata，V。fuscozonatasp。十一月，V. karelica，V. kuusamoensissp。十一月，V。subdevergenssp。nov。，结膜下静脉炎，枯草杆菌和vacillanssp。nov.Verrucaria foveolata在系统发育中嵌套在结膜下V. 但是由于形态和生态地理上的差异，这两个分类单元被视作单独的物种，有待进一步研究。根据分析，研究物种属于Thelidium组。所研究的物种显示出较高的亚种形态，但遗传变异低。此外，它们在形态上显示出相当大的重叠，并且仅基于形态不能可靠地鉴定许多标本。所有种类都限于钙质岩石。Verrucaria alpigena，V。cinereorufa和V. hochstetteri被排除在芬兰的地衣植物区系之外。毛疣疣被认为是一个身份尚未解决的物种。Verrucaria foveolata和V.subtilis在芬兰的钙质岩石上相当普遍，而V. devergens和V.kuusamoensis仅限于芬兰北部。疣膜下疣主要发生在芬兰北部。Verrucaria bifurcata仅在芬兰南部发现。艰难菌Verrucaria difficilis在西南芬兰和东北芬兰都很少。Verrucaria vacillans仅限于芬兰西北角山上的钙质岩石（白云石）。Verrucaria fuscozonata，V。karelica和V. subdevergens仅出现在芬兰东北部的Oulanka地区。为结膜下静脉炎指定一个选型。将芬兰物种的形态与大概属于the族的51种欧洲疣疣物种进行了比较。Verrucaria fuscozonata，V。karelica和V. subdevergens仅出现在芬兰东北部的Oulanka地区。为结膜下静脉炎指定一个选型。将芬兰物种的形态与大概属于the族的51种欧洲疣疣物种进行了比较。Verrucaria fuscozonata，V。karelica和V. subdevergens仅出现在芬兰东北部的Oulanka地区。为结膜下静脉炎指定一个选型。将芬兰物种的形态与大概属于the族的51种欧洲疣疣物种进行了比较。