The genus Scyliorhinus is part of the family Scyliorhinidae, the most diverse family of sharks and of the subfamily Scyliorhininae along with Cephaloscyllium and Poroderma. This study reviews the phylogenetic relationships of species of Scyliorhinus in the subfamily Scyliorhininae. Specimens of all Scyliorhinus species were examined as well as specimens of four of the 18 species of Cephaloscyllium, two species of Poroderma, representatives of almost all other catshark (scyliorhinid) genera and one proscylliid (Proscyllium habereri). A detailed morphological study, including external and internal morphology, morphometry and meristic data, was performed. From this study, a total of 84 morphological characters were compiled into a data matrix. Parsimony analysis was employed to generate hypotheses of phylogenetic relationships using the TNT 1.1. Proscyllium habereri was used to root the cladogram. The phylogenetic analysis, based on implied weighting (k = 3; 300 replications and 100 trees saved per replication), resulted in three equally most parsimonious cladograms with 233 steps, with a CI of 0.37 and an RI of 0.69. The monophyly of the subfamily Scyliorhininae is supported as well as of the genus Scyliorhinus, which is proposed to be the sister group of Cephaloscyllium. The phylogenetic relationships amongst Scyliorhinus species are presented for the first time.

中文翻译：

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基于比较形态学的 Scyliorhinus 属 (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) 猫鲨的系统发育关系

Scyliorhinus 属是 Scyliorhinidae 家族的一部分，Scyliorhinidae 是最多样化的鲨鱼家族，与 Cephaloscyllium 和 Poroderma 一起属于 Scyliorhininae 亚科。本研究回顾了 Scyliorhininae 亚科中 Scyliorhinus 物种的系统发育关系。检查了所有 Scyliorhinus 物种的标本以及 18 种 Cephaloscyllium 中的四种、两种 Poroderma、几乎所有其他猫鲨 (scyliorhinid) 属的代表和一种 proscylliid (Proscyllium habereri) 的标本。进行了详细的形态学研究，包括外部和内部形态、形态测量和分生数据。从这项研究中，总共 84 个形态特征被编译成一个数据矩阵。使用简约分析来生成使用 TNT 1.1 的系统发育关系假设。Proscyllium habereri 被用来为分支图生根。系统发育分析基于隐含权重（k = 3；300 次重复，每次重复保存 100 棵树），产生三个同样最简约的分支图，具有 233 个步骤，CI 为 0.37，RI 为 0.69。支持 Scyliorhininae 亚科和 Scyliorhinus 属的单系，后者被提议为 Cephaloscyllium 的姐妹群。首次提出了 Scyliorhinus 物种之间的系统发育关系。支持 Scyliorhininae 亚科和 Scyliorhinus 属的单系，后者被提议为 Cephaloscyllium 的姐妹群。首次提出了 Scyliorhinus 物种之间的系统发育关系。支持 Scyliorhininae 亚科和 Scyliorhinus 属的单系，后者被提议为 Cephaloscyllium 的姐妹群。首次提出了 Scyliorhinus 物种之间的系统发育关系。