The genus Lodderena Iredale, 1924 has been classified in the Skeneidae by most recent authors. However, this family, originally characterized by their minute size, lack of nacre in the teleoconch and a rhipidoglossate radula, is currently considered to be polyphyletic assemblage, and preliminary molecular systematic data suggest exclusion of Lodderena from Skeneidae. In order to shed light on the systematic position of this genus, we provide a detailed description of the anatomy and histology of the type species, Lodderena minima (Tenison-Woods, 1878), and of L. ornata (Olsson & McGinty, 1958). The anatomical data confirm the vetigastropod-trochoid nature of Lodderena but exclude the genus from Skeneidae. Skeneidae are mainly characterized by a propodial penis, a hermaphroditic condition with separated testis and ovary, and a large receptaculum in the mantle roof. In contrast, Lodderena species lack both a copulatory organ and receptaculum, and have a true ovotestis. We also provide a critical review of nominal species in the genus. Based on molecular and morphological evidence, we exclude Lodderena from Skeneidae, instead treating it as a genus incertae sedis within Trochoidea. We discuss the implications of small size for functional morphology and reproduction.

中文翻译：

---

Skeneimorph而不是Skeneid：Lodderena Iredale的系统学和解剖学，1924年（Vetigastropoda：Trochoidea）

最近的作者将1924年的Lodderena Iredale属分类为Skeneidae。然而，该家族最初以其微小的大小，远距海螺中缺少珍珠质和根状光滑的小舌状茎为特征，目前被认为是多系组合，并且初步的分子系统数据表明，罗氏假单胞菌被排除在Skeneidae外。为了阐明该属的系统位置，我们详细描述了小种Lodderena minima（Tenison-Woods，1878）和L. ornata（Olsson＆McGinty，1958）的解剖学和组织学。。解剖学数据证实了罗德雷纳（Lodderena）的天体摆线性但不包括Skeneidae的属。Skeneidae的主要特征是阴茎发育不佳，睾丸和卵巢分开的雌雄同体状态以及在地幔顶上的大型接受器。相比之下，罗德勒纳犬种既缺乏交配器官，又缺乏受体，并且具有真正的卵睾丸。我们还对属中的标称物种进行了严格的审查。基于分子和形态学证据，我们排除Lodderena从Skeneidae，而不是把它当作属地位未定内斯氏藻。我们讨论了小尺寸对功能形态和繁殖的影响。