Abstract Zuberia is a leaf genus of the Mesozoic seed fern foliage that belongs to Corystospermales. Dispersed leaf cuticles of Zuberia papillata are described using conventional light and electron microscopy techniques (LM, SEM, and TEM). The cuticles were found in Hoyada de Ischigualasto, Ischigualasto Formation (Early Neotriassic), San Juan Province, Argentina. This research provides a new approach that may be useful for the comparison of the cuticle fine details among Zuberia species and other pteridosperms. The cuticle shows characters such as thick cuticle, papillae, stomatal apparatuses sunken and subsidiary cells with papillae overarching the suprastomatal aperture, which suggests adaptations to a megamonsoonal seasonal climatic regime as proposed to the Ischigualasto Formation. Furthermore, cuticle ultrastructural characters reveal new data about the paleoecology and taxonomic status of Zuberia.

中文翻译：

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阿根廷圣胡安省 Hoyada de Ischigualasto（上三叠纪）Zuberia papillata（Townrow）Artabe 1990 叶角质层精细结构的研究

摘要 Zuberia 是属于Corystospermales 的中生代种子蕨叶的叶属。使用传统的光学和电子显微镜技术（LM、SEM 和 TEM）描述了 Zuberia papillata 的分散叶角质层。在阿根廷圣胡安省伊斯基瓜拉斯托组（新三叠纪早期）的霍亚达德伊斯基瓜拉斯托发现了这些角质层。这项研究提供了一种新方法，可能有助于比较 Zuberia 物种和其他蕨类植物的角质层细节。角质层显示出诸如厚角质层、乳头、凹陷的气孔器和带有乳头的附属细胞等特征，乳头覆盖着气孔上孔，这表明适应了对 Ischigualasto 组提出的巨型季风季节性气候制度。此外，