Interpretation of the floral structure of Zingiberaceae has long concentrated on the relationships of the androecial members. It suggested that labellum is composed of two structures rather than three or five, and glands are interpreted either as gynoecial part or as androecial members. Serial sections were used to observe the vasculature of normal and two-staminate flowers in Alpinia intermedia ‘shengzhen’. Floral diagrams were drawn to interpret the morphological category of the floral organs and the relationships of the androecial members. Androecial vascular bundles were associated with carpellary dorsal bundles (CDBs) and parietal bundles (PBs) in a Zingiberales phylogeny setting using ancestral state reconstruction. Anatomical observations demonstrate that the fertile stamen(s) incorporate parietal bundles both in normal and two-staminate flowers. The three appendages represent the three members of the outer whorl of the androecium, while the labellum represents the inner whorl of the androecium in the two-staminate flower. Reconstruction of the origin of the vascular system in the androecium suggests that the outer whorl of androecium receives its vascular supply from the CDBs, and the inner whorl of androecium receives from the PBs in both the basal banana group and the more derived ginger clade. The present study adds to a growing body of literature suggesting that the anatomy of abnormal flowers may not provide enough evidence for elucidating the relationships of the androecial members, and help us to better understand how the vascular system is constructed during the androecial petaloidy evolution.

中文翻译：

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异常花朵的解剖结构能否阐明姜（姜科）中雄性成员的关系？

姜科花结构的解释长期以来一直集中在雄蕊成员的关系上。它表明唇唇由两个结构组成，而不是三个或五个结构，腺体被解释为女性部分或男性部分。采用连续切片观察高良姜'盛珍'中正常花和二雄花的脉管系统。绘制花图以解释花器官的形态类别和雄蕊成员的关系。在使用祖先状态重建的姜目系统发育环境中，雄蕊维管束与心皮背束 (CDBs) 和顶叶束 (PBs) 相关。解剖观察表明，可育雄蕊在正常和双雄花中都包含顶叶束。三个附属物代表雄蕊外轮的三个成员，而唇瓣代表两雄花中雄蕊的内轮。雄蕊血管系统起源的重建表明雄蕊的外轮从CDBs获得血管供应，雄蕊的内轮从基础香蕉组和更衍生的姜分支中的PBs获得血管供应。本研究增加了越来越多的文献，表明异常花朵的解剖结构可能无法提供足够的证据来阐明雄性成员的关系，