While contemplating what to write about my late colleague Denis, whose many outstanding contributions have left an everlasting fingerprint on our knowledge of Great Lakes ciliates, foodwebs and general ecology, I had a sudden leap down memory lane. It all started when our phycologist observed a considerable number of ciliates while analyzing Lake Ontario phytoplankton samples in the early 1990s. Initially, we did not pay much attention to those ciliates, focusing instead on algal particles. However, we came across so many of them that I contacted Dr. William Taylor, a professor at the nearby University of Waterloo, for advice. I knew Bill as a specialist in ciliates and zooplankton. Bill assisted us with the initial identifications and counting, but when the task grew beyond our expectations he recommended that we contact Denis Lynn at the University of Guelph. Denis showed interest in the project but was very frank in indicating that he had no previous experience with Great Lakes ciliates. This was undoubtedly true, since very little was known about the ciliates of the Great Lakes at that time. Luckily for us, Denis agreed to analyze Lake Ontario samples, with a small contract for a trained student under his supervision. Thus began our collaboration, which continued well after his retirement from Guelph in 2010. Even now, we continue to collaborate with Denis’s lab through the excellent support of his long-time associate, Michaela Str€uder-Kypke. For this anecdotal piece, I started to review Denis’s co-operation with our lab, which included almost three decades’ worth of collaborations with Denis on foodweb ecology in the Great Lakes. From 1990 to 2018 our collaborations were very productive, with 19 conference presentations, five book chapters, and six primary publications (Table 1a,b). The presentations show a variety of ciliate data collected in individual ecosystems ranging from Lakes Ontario, Erie, Huron, and Superior to the Bay of Quinte, as well as a comparison of all the Great Lakes. The 11 peer-reviewed publications similarly include a range of coverage sequentially from Lakes Erie, Huron, Ontario, Michigan, and Superior, the Bay of Quinte, and Hamilton Harbour, with comparisons of all the Great Lakes. Some noteworthy publications include Lynn and Munawar, 1999; Munawar and Lynn, 2002; Lynn et al., 2003; and Munawar et al., 2013, which provide first-time insight into the abundance, composition, grazing characteristics, and ecology of the Great Lakes ciliates and their interactions in the microbial loop. On the social side, I remember Denis as a likeable, charming, and outgoing man. During one of his visits to our laboratory in Burlington, we came to know that he liked international food, including Indian cuisine. Consequently, I was pleased to take him for an Indian lunch in one of the local restaurants, where he greatly enjoyed the tasty dishes. As a protistologist, Denis should be credited with creating a unique database of ciliates in the Great Lakes in a relatively short period. Fisheries and Oceans Canada recognized the significant

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与纤毛虫大师合作：与丹尼斯林恩在五大湖的合作和成就

在考虑如何写我已故同事丹尼斯的文章时，他的许多杰出贡献在我们对五大湖纤毛虫、食物网和一般生态学的知识中留下了永恒的印记，我突然想起了回忆。这一切都始于 1990 年代初，我们的藻类学家在分析安大略湖浮游植物样本时观察到大量纤毛虫。最初，我们并没有过多关注那些纤毛虫，而是关注藻类颗粒。然而，我们遇到了很多这样的问题，以至于我联系了附近滑铁卢大学的教授威廉·泰勒博士寻求建议。我认识比尔是纤毛虫和浮游动物的专家。比尔协助我们进行初步鉴定和计数，但是当任务超出我们的预期时，他​​建议我们联系圭尔夫大学的丹尼斯林恩。丹尼斯对该项目表现出兴趣，但非常坦率地表示他之前没有接触过五大湖纤毛虫。这无疑是正确的，因为当时对五大湖的纤毛虫知之甚少。对我们来说幸运的是，丹尼斯同意分析安大略湖的样本，并在他的监督下与一名受过训练的学生签订了一份小合同。我们的合作就这样开始了，在他 2010 年从圭尔夫退休后继续保持良好的合作关系。即使是现在，在丹尼斯的长期合作伙伴 Michaela Str€uder-Kypke 的大力支持下，我们仍在继续与他的实验室合作。对于这个轶事，我开始回顾丹尼斯与我们实验室的合作，其中包括与丹尼斯在五大湖的食物网生态学方面进行了近三年的合作。从 1990 年到 2018 年，我们的合作非常富有成效，有 19 场会议报告、5 本书章节和 6 篇主要出版物（表 1a、b）。演示文稿展示了从安大略湖、伊利湖、休伦湖和苏必利尔湖到昆特湾的各个生态系统中收集的各种纤毛虫数据，并对所有五大湖进行了比较。11 份经过同行评审的出版物同样涵盖了伊利湖、休伦湖、安大略湖、密歇根湖和苏必利尔湖、昆特湾和汉密尔顿港的一系列报道，并对所有五大湖进行了比较。一些值得注意的出版物包括 Lynn 和 Munawar，1999 年；穆纳瓦尔和林恩，2002 年；林恩等人，2003 年；和 Munawar 等人，2013 年，首次了解五大湖纤毛虫的丰度、组成、放牧特征和生态以及它们在微生物循环中的相互作用。在社交方面，我记得丹尼斯是一个讨人喜欢、迷人且外向的人。在他访问我们位于伯灵顿的实验室期间，我们了解到他喜欢国际美食，包括印度美食。因此，我很高兴带他去当地一家餐馆吃印度午餐，他非常喜欢那里的美味佳肴。作为一名原生生物学家，丹尼斯应该在相对较短的时间内创建了五大湖区纤毛虫的独特数据库。加拿大渔业和海洋部认识到重要的 五大湖纤毛虫的生态学及其在微生物循环中的相互作用。在社交方面，我记得丹尼斯是一个讨人喜欢、迷人且外向的人。在他访问我们位于伯灵顿的实验室期间，我们了解到他喜欢国际美食，包括印度美食。因此，我很高兴带他去当地一家餐馆吃印度午餐，他非常喜欢那里的美味佳肴。作为一名原生生物学家，丹尼斯应该在相对较短的时间内创建了五大湖区纤毛虫的独特数据库。加拿大渔业和海洋部认识到重要的 五大湖纤毛虫的生态学及其在微生物循环中的相互作用。在社交方面，我记得丹尼斯是一个讨人喜欢、迷人且外向的人。在他访问我们位于伯灵顿的实验室期间，我们了解到他喜欢国际美食，包括印度美食。因此，我很高兴带他去当地一家餐馆吃印度午餐，他非常喜欢那里的美味佳肴。作为一名原生生物学家，丹尼斯应该在相对较短的时间内创建了五大湖区纤毛虫的独特数据库。加拿大渔业和海洋部认识到重要的 因此，我很高兴带他去当地一家餐馆吃印度午餐，他非常喜欢那里的美味佳肴。作为一名原生生物学家，丹尼斯应该在相对较短的时间内创建了五大湖区纤毛虫的独特数据库。加拿大渔业和海洋部认识到重要的 因此，我很高兴带他去当地一家餐馆吃印度午餐，他非常喜欢那里的美味佳肴。作为一名原生生物学家，丹尼斯应该在相对较短的时间内创建了五大湖纤毛虫的独特数据库。加拿大渔业和海洋部认识到重要的