ABSTRACT Introduction. Within Racomitrium Sect. Racomitrium, several species are distinguished in the Southern Hemisphere, but only the variable R. lanuginosum (Hedw.) Brid. exists in the north. Plants deviating from the common R. lanuginosum phenotype in leaf orientation and in almost entire or weakly dentate hair-point margins suggest that more than one species may exist in this region. Methods. Sixty-five specimens of R. lanuginosum, including eleven deviating ones (four growing intermixed with ‘normal’ plants), and two of each of the Southern Hemisphere R. geronticum Müll.Hal. and R. pruinosum (Wilson) Müll.Hal. were studied, based on the nuclear ITS and the plastid rpl16 and trnG. Relationships among specimens were explored with NeighborNet split networks, maximum parsimony, and Jacknife analyses. Key results. The molecular analyses revealed one grade and four distinct lineages, two of which correspond with the Southern Hemisphere species. Two of the three R. lanuginosum entities are widespread in Scandinavia, whereas the third occurs only in the mountains. Morphologically deviating plants do not form their own lineage but appeared in all three R. lanuginosum entities. Conclusions. The three entities within R. lanuginosum do not form a monophyletic group, good support exists for their recognition, and they are molecularly as distinct as the two morphologically recognisable Southern Hemisphere species. They should therefore be recognised as cryptic species. The morphologically deviating plants represent genotypic differentiation, which was further confirmed by molecular evidence in three out of four occurrences where they grew intermixed with ‘normal’ plants. Since deviating plants occur in all three cryptic species they should not be taxonomically recognised.

中文翻译：

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斯堪的纳维亚 Racomitrium lanuginosum (Hedw.) Brid 中揭示的隐秘物种形成。（菊科）

摘要介绍。在 Racomitrium 教派内。Racomitrium，几个物种在南半球有区别，但只有可变 R. lanuginosum (Hedw.) Brid。存在于北方。在叶方向和几乎整个或弱齿状毛点边缘偏离常见的 R. lanuginosum 表型的植物表明，该地区可能存在不止一种物种。方法。65 个 R. lanuginosum 标本，包括 11 个变异标本（四个与“正常”植物混合生长），以及南半球 R. geronticum Müll.Hal 各两个。和 R. pruinosum (Wilson) Müll.Hal。研究了基于核 ITS 和质体 rpl16 和 trnG。使用 NeighborNet 分裂网络、最大简约和 Jacknife 分析探索了样本之间的关系。关键结果。分子分析揭示了一个等级和四个不同的谱系，其中两个与南半球物种相对应。三个 R. lanuginosum 实体中有两个广泛分布在斯堪的纳维亚半岛，而第三个仅出现在山区。形态上偏离的植物不会形成它们自己的谱系，而是出现在所有三个 R. lanuginosum 实体中。结论。R. lanuginosum 中的三个实体不形成单系群，它们的识别存在良好的支持，并且它们在分子上与两种形态上可识别的南半球物种一样不同。因此，它们应该被认为是神秘物种。形态上偏离的植物代表基因型分化，在四分之三的事件中，它们与“正常”植物混合生长，分子证据进一步证实了这一点。由于变异植物出现在所有三个隐秘物种中，因此它们不应在分类学上得到认可。