Cafius gigas Lea, 1929 (Coleoptera: Staphylinidae) was a large rove beetle endemic to Lord Howe Island (LHI) resembling Cafius and the LHI flightless endemic Hesperus dolichoderes (Lea, 1925). Like several other LHI endemics, C. gigas became extinct due to human-introduced rats. It is a legacy species valuable for understanding the LHI biota in terms of evolutionary biology and historical biogeography. Whether C. gigas was a member of Cafius Curtis, 1829, restricted to oceanic shores and prone to trans-oceanic dispersal, or related to H. dolichoderes, would have different implications. We subjected C. gigas to a total-evidence phylogenetic analyses of morphological and molecular data using model-based and parsimony methods. As a result, it is transferred to Hesperus Fauvel, 1874 with the new combination Hesperus gigas (Lea, 1929) comb. nov. Our analysis indicates that the montane leaf litter inhabitant H. gigas evolved neither in situ nor from a seashore Cafius-ancestor, or from an ancestor shared by two other LHI endemic congeners, Hesperus pacificus Olliff, 1887 and H. dolichoderes. It also suggests that all three Hesperus species that currently occur on LHI could have evolved on various seamounts at various times before reaching LHI.

中文翻译：

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全面证据的方法解决了Cafius gigas的系统发育问题，这是豪勋爵岛上最近灭绝的独特甲虫

Cafius gigas Lea，1929年（鞘翅目：Staphylinidae）是豪勋爵岛（LHI）的大型ro虫，类似于Cafius和LHI不能飞行的地方性Hesperus dolichoderes（Lea，1925年）。像其他几种LHI特有的一样，由于人类引入的老鼠，C。gigas灭绝了。它是一个遗产物种，对于了解LHI生物区系的进化生物学和历史生物地理学非常有价值。吉格斯犬（C. gigas）是否是1829年的卡菲乌斯·柯蒂斯（Cafius Curtis ）的一员，只限于大洋沿岸并易于越洋扩散，还是与多毛线虫相关，所以具有不同的含义。我们接受了C. gigas使用基于模型和简约的方法对形态和分子数据进行系统证据的系统发育分析。结果，它以新的组合Hesperus gigas（Lea，1929）梳子转移到Hesperus Fauvel，1874年。十一月 我们的分析表明，山地落叶凋落物H. gigas既不是就地进化也不是从海滨Cafius-祖先进化而来，也不是由LHI的两个其他本地同类人Hesperus pacificus Olliff（1887）和H. dolichoderes共同进化而成。这也表明所有三个Hesperus 目前在LHI上发生的物种在到达LHI之前可能已经在不同的海山上演化。