Abstract

The structure of Thymus petraeus individuals has been studied architecturally. It has been established that in the south of Siberia the species occurs in similar habitats: in true, meadow, sandy steppes and their petrophytic variants on the slopes and low rocky peaks of hills. According to the classification proposed by I.G. Serebryakov (1964), T. petraeus is a vegetatively semimobile dwarf shrub with a system of the main root that is persistent throughout its life and with rooting runners. Three architectural units that differ in the spatial position of the compound skeletal axis (orthotropic–plagiotropic, plagiotropic, and orthotropic) have been identified in the structure of T. petraeus individuals. Each architectural unit consists of the main compound skeletal axis, compound skeletal axes of the 1st order, formation shoots, branching shoots, and enrichment shoots. It is shown that the structure of individuals can be formed by repeating only two architectural units. It is established that the structure of individuals growing in the true steppes and their petrophytic variants is built at the expense of the repetition of orthotropic–plagiotropic and plagiotropic architectural units. That in sandy steppes is built at the expense of repetition of plagiotropic and orthotropic architectural units. The features of T. petraeus development in specific ecological and coenotic conditions are shown. In this regard, we have characterized (1) morphological polyvariance, which is based on changes in the structure of shoots (shortened and elongated) and the composition of architectural units (no compound skeletal axes of the 1st order or enrichment shoots); (2) dimensional polyvariance, manifested in a change in the length and number of compound skeletal axes; and (3) dynamic polyvariance, associated with fluctuations in the duration of the monopodial growth of shoots formation (from 2 to 5 years) and the architectural unit as a whole (from 6 to 20 years). The revealed modifications of architecture do not lead to a change in the life form of the dwarf shrub, but reflect the mechanisms of its adaptation.

中文翻译：

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西伯利亚南部条件下矮灌木胸腺petraeus（唇形科）的体系结构

摘要

百里香（Thymus petraeus）个体的结构已在建筑上进行了研究。已经确定，该物种在西伯利亚南部出现在相似的栖息地：在真实的草甸，沙质草原及其山坡和岩石低峰的岩石变体中。根据IG Serebryakov（1964）提出的分类，T。petraeus是一种植物性半流动性矮化灌木，其主要根系贯穿其一生，并带有生根赛跑者。已在T. petraeus的结构中确定了三个在复合骨骼轴的空间位置不同的建筑单元（正交各向异性-斜向性，斜向性和正交性）个人。每个建筑单元均由主要的复合骨架轴，一阶复合骨架轴，地层芽，分支芽和富集芽组成。结果表明，仅通过重复两个建筑单元就可以形成个体的结构。可以确定的是，在真正的草原上生长的个体及其岩石质变体的结构是建立在正交异性-斜生性和斜生性建筑单元重复的基础上的。在沙质草原中建造的建筑是以重复造斜和正交各向异性的建筑单元为代价的。彼德罗非鱼的特征显示了在特定的生态和共生条件下的发育。在这方面，我们已经表征（1）形态多方差，它是基于枝条结构（变短和延长）和建筑单元组成的变化（没有一阶的复合骨架轴或富集枝条）引起的；（2）尺寸多方差，表现为复合骨骼轴的长度和数量的变化；（3）动态多方差，与芽形成单足类生长的持续时间（2至5年）和整个建筑单位（6至20年）相关。所揭示的建筑修改不会导致矮灌木的生活形式发生变化，而是反映了其适应机制。