The group Tapejaridae forms a clade of toothless pterosaurs easily recognized by their premaxillary sagittal crests and particularly large nasoantorbital fenestrae. The tapejarids represent the most representative group of pterosaurs from the Lower Cretaceous Crato Formation of the Araripe Basin (Northeastern Brazil). The holotype of the large tapejarid Tupandactylus imperator Campos and Kellner, 1997 is known by two main slabs from the New Olinda Member of the Crato Formation, however, only one of the slabs containing the sagittally bipartite skull is referred to the holotype of Tupandactylus imperator, remain the counter-slab be properly described. The cotype is fragmented in several broken pieces and presents a significative number of cranial elements. A medial internasal septum completely preserved inside the nasoantorbital fenestra is reported for the first time for pterosaurs. The exceptional preservation of a collagenous septum and other integumentary structures visible in the cotype specimen is extremely rare and supports the concept of the unusual pattern of soft tissue observed in the fossils from the Crato Konservat-Lagerstätte, specially pterosaurs. Herein is presented the description of the cotype of Tupandactylus imperator, in complementation to the previously designated slab of the holotype of this tapejarid species. The occurrence of casques in pterosaurs is supported by comparative anatomy with the bird galliform Pauxi (Cracidae). Besides that, it is discussed on the skull with extravagant cranial crests of Tupandactylus imperator and the significance of the associated soft tissues and other cranial integuments, which indicates an expressive morphological and taxonomic diversity among the tapejarid pterosaurs.

中文翻译：

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Crato组的斜齿翼龙Tupandactylus imperator及其颅骨的保存

Tapejaridae组形成无齿翼龙的进化枝，很容易被其上颌前矢状冠和特别是较大的鼻窦门窗齿识别。jar类动物是Araripe盆地（巴西东北部）下白垩统Crato组中最有代表性的翼龙类。大型tape形Tupandactylus imperator的整型由Crato组的New Olinda成员的两个主要平板已知，Campos and Kellner，1997，但是，只有一个包含矢状二分头骨的平板被称为Tupandactylus imperator的整型。，保留对柜台的正确描述。该共型碎片成几个碎片，呈现出大量的颅骨成分。首次报道翼龙内侧鼻中隔完全保留在鼻眶窗内。在共型标本中可见的胶原蛋白隔片和其他外皮结构的特殊保存极为罕见，这支持了在CratoKonservat-Lagerstätte的化石中观察到的软组织异常模式的概念，特别是翼龙。本文介绍了对翼龙的共同类型的描述，以补充此前指定的该类jar类物种的平板。casques在翼龙的发生是由与鸟鸡形目比较解剖学支持Pauxi（凤冠雉科）。除此之外，它还讨论了具霸王龙颅顶的颅骨，以及相关软组织和其他颅骨被膜的重要性，这表明p类翼龙具有表达性的形态学和分类学多样性。