The trace fossil Lumbricaria Münster is a common constituent of the Upper Jurassic lithographic limestones in southern Germany. Despite having been recognized more than 300 years ago, its nature and producer remain enigmatic. Controversial interpretations of Lumbricaria regard it either as the remains of worms, or as cololites (i.e. faecal material preserved in the gut) or coprolites (i.e. ejected faecal material) of worms, fish, holothurians or cephalopods. A morphological analysis of the type material and numerous specimens reveals two basic morphotypes of Lumbricaria, conformable with the ichnospecies L. intestinum (loosely confined, intertwined to elongate) and L. colon (well confined, densely entangled). Transitions and combinations between both ichnospecies exist and are the result of taphonomic effects. A comparison with modern analogues favours cephalopods (in particular nautiloids or ammonites) as the most likely producers of Lumbricaria, an interpretation that is supported by the finding of an Upper Jurassic nautiloid cast preserving a Lumbricaria‐like faecal string within its body chamber. Ammonites are the likely producers of Lumbricaria from the Upper Jurassic of Germany.

中文翻译：

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来自德国上侏罗统的鱼鳞菌（LumbricariaMünster）被解释为粪便中的一串炸药

微量化石LumbricariaMünster是德国南部上侏罗统平纹石灰岩的常见成分。尽管已经有300多年的历史了，但它的性质和制作人仍然是个谜。关于肺mb的有争议的解释是将其视为蠕虫的残留物，还是蠕虫，鱼类，整形人或头足类动物的粪便（即肠道中保存的粪便）或共prolites（即排出的粪便）。该类型的材料和大量标本的形态学分析揭示的两个基本形态Lumbricaria的，适形与ichnospecies L. intestinum（松散地密闭，交织到细长）和L.结肠（密闭，纠缠）。存在两种鱼类物种之间的过渡和组合，这是Thophonomic效应的结果。与现代类似物的比较倾向于将头足类（特别是鹦鹉螺或or虫）作为mb类的最可能的生产者，这一解释得到了上侏罗统鹦鹉螺类铸件的发现，该铸件在其体腔内保留了类似mb类的粪便串。炸药很可能是德国上侏罗统的腰mb的生产者。