Heteropterans communicate chemically through thoracic or abdominal glandular complexes. The dorso-abdominal scent glands (DAGs) are externalized by cuticular specialized structures forming the external scent efferent system (SES). Different groups in Heteroptera present other glands, such as the Asopinae (Pentatomidae), whose males can have ventral abdominal pheromonal glands externalized by cuticular modifications, called glandular patches (GPs). Here we describe the SES of DAGs and, for the first time, the GPs of 22 Asopinae genera. The ostioles of SES 1 vary in shape. SES 2 and SES 3 were restricted to scars, indicating that only the SES 1 remains functional in adults. The GPs are found mostly in segments V and VI. We have not found any difference in the morphological pattern between the SES of species bearing GPs and those lacking it, meaning that the acquisition of GPs is likely related to some sexual behavior, thus not as a complex structure replacing any role of the DAGs. The cuticle of the GPs is microsculptured, bearing many pores surrounding the base of sensilla being the pathway for the secretions to reach the outside of the exoskeleton. The sensilla may be adaptations to increase the contact surface, assisting rapid evaporation of the volatile secretions from the pores.

中文翻译：

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腹足类（半翅目：异翅类：五翅目）腹腺的外部形态。

异翅类动物通过胸或腹腺复合体进行化学通讯。背腹气味腺（DAG）由形成外部气味传出系统（SES）的表皮专门结构外部化。异翅目中的不同群体还存在其他腺体，例如Asopinae（Pentatomidae），它们的雄性可以具有被表皮修饰外化的腹腹腹侧腺体，称为腺体贴片（GPs）。在这里，我们描述了DAG的SES，并且首次描述了22个Asopinae属的GP。SES 1的口骨形状各异。SES 2和SES 3仅限于疤痕，这表明只有SES 1在成年人中仍然可以发挥功能。GP大多位于V和VI段中。我们还没有发现带有GP的物种和缺乏GP的物种的SES在形态学模式上没有任何区别，这意味着，获得全科医生可能与某些性行为有关，因此不是取代DAG的任何角色的复杂结构。GP的角质层是经过微雕的，在感官基部周围有许多孔，这些孔是分泌物到达外骨骼外部的途径。感觉器可以是适应性的，以增加接触表面，从而帮助从孔中快速蒸发出挥发性分泌物。