Analysis of mushroom‐shaped rugose corals Schlotheimophyllum patellatum (Schlotheim, 1820) from the Silurian (Upper Visby Beds, Lower Wenlock, Sheinwoodian) of Gotland, Sweden, showed that they were colonized on both the upper (exposed) and lower (cryptic) sides by a variety of encrusting and boring (sclerobiont) biotas, represented by 10 taxa and at least 23 species. Bryozoans and microconchid tubeworms, the most abundant encrusters, dominated on the cryptic undersides of the corals, while the dominant endobionts responsible for Trypanites borings overwhelmingly dominated the exposed surfaces. Except for cnidarian sphenothallids, which were exclusive colonizers of the underside of only one coral host, no other encrusters could be referred to as obligate cryptobionts. Because the upper surface of these corals was likely covered by soft‐tissues during life, in specimens lifted off the sea‐floor sclerobionts must have settled on the cryptic sides first. They could colonize the upper side only after the coral’s death, unless it was covered by sediment as could be the case in some flat specimens. With time, the space on the underside of the coral skeleton may have progressively been filled by sediment as well, precluding further colonization by sclerobionts. In that respect, the colonization patterns of these corals by encrusters and borers were controlled by the complex interplay of environmental factors, sclerobiont dynamics and coral growth in a given Silurian habitat. Compared with Silurian stromatoporoid hosts, the sclerobiont diversity and abundance noted on the Schlotheimophyllum corals may be regarded as representative for the Silurian as a whole.

中文翻译：

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瑞典哥得兰岛志留纪的皱纹珊瑚Schlotheimophyllum patellatum（Schlotheim，1820）上的硬核菌定殖模式

对来自瑞典哥得兰岛的志留系（上维斯比河床，下温洛克，希恩伍德人）的蘑菇状皱纹珊瑚Schlotheimophyllum patellatum（Schlotheim，1820）进行分析，结果表明它们在上（裸露）和下（隐蔽）侧均被定殖由各种各样的包缠和无聊的生物组成，以10个分类单元和至少23种为代表。苔藓虫和微孔虫是最丰富的结壳动物，在珊瑚的隐秘底面占主导地位，而负责锥虫的主要内生生物钻孔绝大多数占据了裸露的表面。除了CNIDIAN Sphenothallids是仅一个珊瑚宿主下侧的独家定居者外，其他包壳都不能被称为专性隐孢子虫。由于这些珊瑚的上层在生活中很可能被软组织覆盖，因此从海底举起的硬骨鱼标本必须首先安放在隐秘的一面。他们只有在珊瑚死亡后才能在上侧殖民，除非像某些平坦的标本一样被沉积物覆盖。随着时间的流逝，珊瑚骨骼下面的空间可能也逐渐被沉积物填充，从而阻止了硬骨s菌的进一步定居。在这方面，在给定的志留纪生境中，环境因素，硬皮病动态和珊瑚生长之间复杂的相互作用控制着由甲壳虫和蛀虫对这些珊瑚的定殖模式。与志留纪层间质多孔宿主相比，石笋的多样性和丰度在Schlotheimophyllum珊瑚可以看作是志留纪整体的代表。