The palaeobiogeography of the alveolinoid Borelis species reveals the evolutionary patterns leading to the two extant representatives, which occur in shallow‐water tropical carbonate, coral reef‐related settings. Type material and new material of fossil Borelis species, along with Recent specimens were studied to assess their taxonomic status, species circumscriptions (based on proloculus size, occurrence of Y‐shaped septula, and the index of elongation), palaeobiogeography and evolutionary dynamics. The species dealt with here are known from exclusively fossil (B. pygmaea, B. inflata, B. philippinensis, B. melo, B. curdica), and from fossil and modern (B. pulchra, B. schlumbergeri) specimens. For the first time, fossil and Recent Borelis specimens are illustrated via micro‐computed tomography scanning images. Depending on the occurrence of Y‐shaped septula, two lineages are distinguished. Deriving from the middle–upper Eocene Borelis vonderschmitti, the first lineage includes B. inflata, B. pulchra and B. pygmaea, lacking Y‐shaped septula. The first species bearing Y‐shaped septula is the Rupelian B. philippinensis of the western Indo‐Pacific. The westward migrants of B. philippinensis into the Mediterranean gave rise to B. melo (Aquitanian–Messinian) and B. curdica (Burdigalian–Tortonian). These two species became isolated from the Indo‐Pacific by the Langhian eastern closure of the Mediterranean basin and disappeared during the Messinian Salinity Crisis. Since the Tortonian, B. schlumbergeri, which descended from B. philippinensis, has inhabited the Indo‐Pacific along with B. pulchra. From the central Pacific Ocean, B. pulchra reached the Caribbean area before the early Piacenzian closure of the Central America seaway.

中文翻译：

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较大的有孔虫Borelis de Montfort（Borelidae）的古生物地理学和演化模式

肺泡类Borelis物种的古生物地理学揭示了导致这两个现存代表的演化模式，它们发生在浅水热带碳酸盐岩，与珊瑚礁相关的环境中。研究了化石Borelis物种的类型材料和新材料，以及最近的标本，以评估它们的分类学地位，物种限制（基于前lo大小，Y形中隔的出现以及伸长指数），古生物地理学和进化动力学。该物种在这里讨论从化石专用（被称为B.矮，  B.胀，  B. philippinensis，  B.甜瓜，  B. curdica），并从化石和现代（B. pulchra，  B。schlumbergeri）标本。通过显微计算机断层扫描图像首次展示了化石和最近的Borelis标本。根据Y形间隔的出现，可以区分两个谱系。起源于中新世始新世Borelis vonderschmitti，第一个世系包括B. inflata，  B. pulchra和B. pygmaea，缺少Y形的中隔。第一个带有Y型分隔符的物种是印度洋西部的Rupelian B. philippinensis。B. philippinensis的向西迁移到地中海引起了B. melo（阿基坦人-麦西尼亚人）和B. curdica（布迪加利安人-托托尼亚人）。这两个物种通过地中海盆地的Langhian东区封闭而与印度太平洋隔离开来，并在Mesinian盐度危机期间消失了。自Tortonian以来，源自philippinensis的B. schlumbergeri便与B. pulchra一起生活在印度太平洋。B. pulchra从中部太平洋到达加勒比海地区，之后中美洲海道的Piacenzian早期关闭。