A new fossil batoid (ray) Ptychotrygon rostrispatula sp. nov. is described from five exceptionally well‐preserved, three‐dimensional skeletal remains from the Turonian (Late Cretaceous) of Morocco. These specimens represent the first known skeletal remains for the genus Ptychotrygon and allow an almost complete description of the genus, providing a new insight into its phylogenetic relations and validating its taxonomic status as a member of the Sclerorhynchoidei. Mechanical preparation of the fossil remains revealed a relatively large batoid species (estimated total length above 1 m). Overall the morphology resembles that of sclerorhynchoids, with a robust hypertrophied rostrum that lacks enlarged rostral denticles with enlarged paddle‐like pectoral proximal elements (propterygium, mesopterygium and metapterygium). Never before seen details of the branchial skeleton are presented (large second hypobranchial without anterior process, which was probably fused to the basibranchial and no evidence of articulation with other branchial element). To assess the phylogenetic relations of these specimens within the sclerorhynchoids a parsimonious analysis using TNT and PAUP was performed. These analyses included Asflapristis cristadentis and Ptychotrygon rostrispatula sp. nov., which were used as representatives of Ptychotrygonidae, and also includes six other genera of sclerorhynchoids with relatively good skeletal remains. Both analyses recovered two monophyletic groups within sclerorhynchoids: the first contains Ischyrhiza, Onchopristis and Schizorhiza, and a second that includes Micropristis, Sclerorhynchus, Libanopristis and Ptychotrygonidae.

中文翻译：

---

摩洛哥Turonian的神秘白垩纪锯try属Ptychotrygon（Chondrichthyes，Batoidea）的第一个骨骼记录

一种新的化石蝙蝠样（射线）Ptychotrygon rostrispatula sp。十一月 它是从摩洛哥的Turonian（晚白垩世）时期保存的五个保存最完好的三维骨骼遗骸中描述的。这些标本代表了Ptychotrygon属的第一个已知骨骼遗骸并可以对该属进行几乎完整的描述，从而为该属的系统发育关系提供新的见解，并验证其作为巩膜螨的一员的分类学地位。机械化石化石残骸的准备工作显示出相对较大的蝙蝠生物（估计总长度在1 m以上）。总体而言，形态类似于巩膜胆管样，具有坚固的肥大的讲台，缺乏增大的延髓表皮齿，以及增大的桨状胸膜近端元件（翼状me肉，中翼状and肉和翼状ery肉）。从未出现过分支骨骼的详细信息（大的第二分支支气管，没有前突，可能与基底支气管融合，没有与其他分支元件咬合的证据）。为了评估这些样本在巩膜扁突内的系统发育关系，使用TNT和PAUP进行了简约分析。这些分析包括Asflapristis cristadentis和Ptychotrygon rostrispatula sp。十一月，被用作Ptychotrygonidae的代表，还包括其他六个具有相对较好的骨骼残留的巩膜扁突属。两种分析回收的2个单系群sclerorhynchoids内：第一个是包含Ischyrhiza，  Onchopristis和Schizorhiza，以及第二包含Micropristis，  Sclerorhynchus，  Libanopristis和Ptychotrygonidae。