The thelypteroid fern genus Stegnogramma s.l. contains around 18–35 species and has a global, cross‐continental distribution ranging from tropical to temperate regions. Several genera and infrageneric sections have been recognized previously in Stegnogramma s.l., but their phylogenetic relationships are still unclear. In this study, we present a global phylogeny of Stegnogramma s.l. with the most comprehensive sampling to date and aim to pinpoint the phylogenetic positions of biogeographically and taxonomically important taxa. Based on the reconstructed historical biogeography and character evolution, we propose a new (infra)generic classification and discuss the diversification of Stegnogramma s.l. in a biogeographical context. New names or combinations are made for 12 (infra)species, including transferring the monotypic species of Craspedosorus to Leptogramma. Finally, we discuss a possible link between leaf architecture and ecological adaptation, and hypothesize that the increase in leaf dissection and free‐vein proportion is an adaptive feature to cool climates in Stegnogramma s.l.

中文翻译：

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Stegnogramma ferns (Thelypteridaceae) 的全球系统发育：一般和部分修订、历史生物地理学和叶结构的进化

Thelypteroid fern 属 Stegnogramma sl 包含大约 18-35 种，具有从热带到温带地区的全球跨大陆分布。先前在 Stegnogramma sl 中已经识别了几个属和属下的部分，但它们的系统发育关系仍不清楚。在这项研究中，我们展示了 Stegnogramma sl 的全球系统发育，具有迄今为止最全面的采样，旨在确定具有生物地理和分类学重要性的分类群的系统发育位置。基于重建的历史生物地理学和特征演变，我们提出了一种新的（下）属分类，并讨论了 Stegnogramma sl 在生物地理学背景下的多样化。为 12 个（以下）物种制作了新名称或组合，包括将 Craspedosorus 的单型物种转移到 Leptogramma。最后，我们讨论了叶片结构与生态适应之间可能存在的联系，并假设叶片解剖和游离叶脉比例的增加是 Stegnogramma sl 对凉爽气候的适应性特征