The brooding gastropod Crepipatella dilatata can feed by scraping the substrate with the radula and by suspension-feeding, which also requires use of the radula. There is a “division of labour” for the radula among three discrete tasks associated with feeding: (1) removing mucous balls from the food pouch; (2) transferring the mucous cord from the neck channel to the mouth (both components of suspension-feeding); (3) scraping the substrate. We hypothesised that the proportion of time used for each feeding activity varies according to environmental conditions. Total radular activity in females was greatest at high tide and in summer. The rate of radular extrusion for ingesting the mucous cord varied seasonally and between brooding and non-brooding females. Non-brooding females exhibited higher rates of radular extrusion for ingesting the mucous cord and for scraping the substrate than did brooders. In females, radular activity in removing the mucous ball from the food pouch was strongly influenced by the tidal cycle during winter, reaching minimum values at low tide. Differences were recorded in substrate scraping among seasons and within tidal cycles, and among males, brooding females and non-brooding females. Brooding females displayed less rasping than non-brooders, since the area available for grazing was restricted by the egg mass. Throughout the year, including low salinity periods, males allocated a greater proportion of total radular activity to rasping than to removing the mucous ball or ingesting the mucous cord. The feeding behaviour of both males and females is modulated by salinity, but the principal determinants of radular activity are the mode of reproduction (brooding in females) and, in males, motility.

中文翻译：

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季节和潮汐周期对河口孵化腹足纲腹足纲腹足feeding（Calippatella dilatata（Calyptraeidae））摄食期间during虫对分工的影响

孵化腹足纲腹足蛾可通过用the刮下基质并通过悬浮进料来进食，悬浮液也需要使用。在与喂养相关的三个不连续的任务中，the的“分工”是：（1）从食品袋中取出粘液球；（2）将粘液从颈部通道转移到口腔（悬浮饲料的两个组成部分）；（3）刮基板。我们假设每次喂养活动所用的时间比例根据环境条件而变化。在涨潮和夏季，雌性的总胚根活动最大。食入粘液的球根挤压的速率随季节变化，在育雏和非育雏雌性之间变化。与育雏者相比，非育雏雌性在吞咽粘液和刮擦基质方面表现出更高的胚根挤压率。在雌性动物中，从食物袋中取出粘液球的球根活动在冬季受到潮汐周期的强烈影响，在退潮时达到最小值。记录的季节和潮汐周期之间以及雄性，育卵雌性和非育卵雌性的底物刮擦差异。育雏的雌性比非育雏的雌性少，因为可放牧的区域受卵量的限制。全年中，包括低盐度时期，雄性将总的胚根活动分配给打磨比分配黏液球或摄取粘液带更多。男性和女性的进食行为受盐度的调节，