Five different ruminant taxa were found in the Gračanica coal mine: the tragulid Dorcatherium vindebonense, a palaeomerycid (Palaeomerycidae gen. et sp. indet.), the giraffe Giraffokeryx sp., and the bovids ?Tethytragus sp. and Eotragus ?clavatus. The ruminant fauna fits well to an early middle to middle middle Miocene age and is typical for a locality of the mammal units MN5 or MN6. The assemblage shows Asian as well as European affinities, represents the westernmost occurrence of Giraffokeryx known so far, and confirms the role of the Balkan Peninsula as a corridor between the two continents. As the ruminant assemblage from Gračanica is not abundant, palaeoecological considerations are only tentative, but the determined taxa point to a predominantly forested environment.

中文翻译：

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来自Gračanica煤矿中新世中期的波斯菊（波斯尼亚和黑塞哥维那布戈尼诺盆地）

在Gračanica煤矿中发现了五种不同的反刍动物分类群：tragulid Dorcatherium v​​indebonense，palaeomerycid（Palaeomerycidae gen。et sp。indet。），长颈鹿Giraffokeryx sp。和bovids 齿螨（Tethytragus sp。）和Eotragus？克拉瓦图斯。反刍动物非常适合中新世中期到中中期，并且对于哺乳动物单位MN5或MN6的位置很典型。集合显示了亚洲和欧洲的相似性，代表了最西部的Giraffokeryx迄今已为人所知，并确认了巴尔干半岛作为两大洲之间走廊的作用。由于来自Gračanica的反刍动物种类不多，因此，古生态学方面的考虑只是暂定的，但确定的分类单元指向主要是森林环境。