The theme of this paper is that Fisher's "Fundamental Theorem of Natural Selection" has various deficiencies as a quantification of the effect of natural selection in a Mendelian population which are not shared by a new different theorem described in this paper. The deficiencies in Fisher's theorem are listed in this paper. The new theorem focuses on the implications of changes in gene frequencies under natural selection and not, as does the Fundamental Theorem of Natural Selection, on changes in mean population fitness. Whereas the algebra in the new theorem corresponds in places to that in the Fundamental Theorem of Natural Selection, the approach, perspective and conclusion of the new theorem are different from those of the Fundamental Theorem of Natural Selection.

中文翻译：

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通过自然选择量化进化。

本文的主题是，费舍尔的“自然选择基本定理”在量化孟德尔人口中自然选择的影响方面存在各种缺陷，而本文所描述的新的不同定理则没有这些缺陷。费舍尔定理的不足在本文中列出。新定理侧重于自然选择下基因频率变化的含义，而自然选择基本定理则不关注平均人口适应性的变化。尽管新定理中的代数与自然选择基本定理在某些地方相对应，但是新定理的方法，观点和结论与自然选择基本定理的方法，观点和结论是不同的。